13F

Stock

Denali Therapeutics Inc.

DNLI · held by 250 managers

Reported value

$2.29B

Holders

250

+2 new · −3 exited

Holders’ est. mark

$15M

Price effect on 10 books still in the name

Largest holder

$215.6M

Vanguard

Notable conviction

Share of each notable manager’s 13F book.

Holder count by quarter

Who holds it

Notable only
InvestorSharesValueEst. markChange
Vanguard13.1M$215.6M—Held
BlackRock, Inc.12.7M$209.9M—Held
BAILLIE GIFFORD & CO12.3M$203.3M—Held
Temasek Holdings (Private) Ltd7.01M$115.8M—Held
State Street6.26M$103.4M—Held
Fidelity6.1M$100.7M—Held
NORGES BANK5.12M$84.5M$8.88MAdded$43.1M
UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC4.98M$82.3M—Held
GOLDMAN SACHS GROUP INC3.84M$63.3M—Held
BAKER BROS. ADVISORS LP3.73M$61.6M—Held
Capital Research Global Investors3.73M$61.6M—Held
ALLIANCEBERNSTEIN L.P.3.56M$58.7M—Held
GEODE CAPITAL MANAGEMENT, LLC3.46M$57.1M—Held
Holocene Advisors, LP3.15M$52.1M—Held
Crestline Management, LP2.81M$46.4M—Held
Fisher Asset Management, LLC2.59M$42.8M—Held
UBS Group AG2.4M$39.7M—Held
DIMENSIONAL FUND ADVISORS LP2.37M$39.2M—Held
Aberdeen Group plc2.03M$33.6M—Held
Casdin Capital, LLC1.84M$30.4M—Held
Lynx1 Capital Management LP1.74M$28.7M—Held
Point72 Asset Management, L.P.1.7M$28.1M—Held
PRINCIPAL FINANCIAL GROUP INC1.52M$25.2M—Held
Siren, L.L.C.1.4M$23.1M—Held
Pictet Asset Management Holding SA1.33M$22M—Held
MORGAN STANLEY1.22M$20.1M$2.43MCut−$4.39M
CHARLES SCHWAB INVESTMENT MANAGEMENT INC1.2M$19.8M—Held
NORTHERN TRUST CORP1.2M$19.8M—Held
Candriam S.C.A.1.17M$19.3M—Held
ExodusPoint Capital Management, LP1.02M$16.9M—Held
Foresite Capital Management VI LLC1M$16.5M—Held
MASTERS CAPITAL MANAGEMENT LLC1M$16.5M—Held
BANK OF AMERICA CORP /DE/932.1K$15.4M$1.85MAdded$1.43M
Alyeska Investment Group, L.P.859.1K$14.2M—Held
Citadel779.1K$12.9M—Held
MARSHALL WACE, LLP763.9K$12.6M—Held
Bank of New York Mellon Corp762.1K$12.6M—Held
Defilade Capital Management, L.P.656.7K$10.8M—Held
Palo Alto Investors LP641.2K$10.6M—Held
Cercano Management LLC625K$10.3M—Held
VOLORIDGE INVESTMENT MANAGEMENT, LLC591.5K$9.77M—Held
Qube Research & Technologies Ltd590.3K$9.75M—Held
SEGALL BRYANT & HAMILL, LLC537K$8.87M—Held
Ensign Peak Advisors, Inc515.7K$8.51M—Held
Vestal Point Capital, LP465K$7.68M—Held
Hudson Bay Capital Management LP450K$7.43M$607KAdded$3M
SCHRODER INVESTMENT MANAGEMENT GROUP451K$7.4M—Held
TWO SIGMA INVESTMENTS, LP430.2K$7.1M—Held
STEMPOINT CAPITAL LP419.1K$6.92M—Held
BARCLAYS PLC407.4K$6.73M$810.6KAdded$380.2K
AMERICAN CENTURY COMPANIES INC379.1K$6.26M—Held
MILLENNIUM MANAGEMENT LLC369.8K$6.1M—Held
Clarius Group, LLC369K$6.09M—Held
TD Asset Management Inc339.4K$5.6M—Held
DAFNA Capital Management LLC318.5K$5.26M—Held
Invesco Ltd.291.9K$4.82M—Held
Marex Group plc287K$4.74M—New
Nuveen, LLC275.5K$4.55M—Held
Swiss National Bank273.8K$4.52M—Held
EDMOND DE ROTHSCHILD HOLDING S.A.225.7K$3.73M—Held
Trexquant Investment LP209.6K$3.46M—Held
BURKEHILL GLOBAL MANAGEMENT, LP200K$3.3M—Held
SCS Capital Management LLC198K$3.27M—Held
WELLS FARGO & COMPANY/MN193.7K$3.2M—Held
Rafferty Asset Management, LLC191.6K$3.16M—Held
MANUFACTURERS LIFE INSURANCE COMPANY, THE184.2K$3.03M—Held
RHUMBLINE ADVISERS183K$3.02M—Held
AMERIPRISE FINANCIAL INC180.2K$2.98M—Held
JPMORGAN CHASE & CO171.9K$2.84M—Held
Susquehanna Portfolio Strategies, LLC167.7K$2.77M—Held
TWO SIGMA ADVISERS, LP154.8K$2.56M—Held
GROUPAMA ASSET MANAGMENT151.6K$2.5M$270.2KAdded$531.6K
Legal & General Group Plc131.2K$2.17M—Held
CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM129.4K$2.14M—Held
MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.128.8K$2.13M—Held
ALPS ADVISORS INC120.7K$1.99M—Held
Woodline Partners LP114.1K$1.88M—Held
B. Metzler seel. Sohn & Co. AG112.3K$1.84M—Held
NOMURA ASSET MANAGEMENT INTERNATIONAL INC.107.5K$1.77M—Held
D.A. DAVIDSON & CO.106.9K$1.77M—Held
DEUTSCHE BANK AG\105.1K$1.73M—Held
Rhenman & Partners Asset Management AB100K$1.65M—Held
Ghisallo Capital Management LLC100K$1.65M—Held
Integrated Wealth Concepts LLC94.6K$1.56M—Held
Y-Intercept (Hong Kong) Ltd93.6K$1.55M—Held
Monashee Investment Management LLC91.6K$1.51M—Held
MetLife Investment Management, LLC75.2K$1.24M—Held
STATE OF WISCONSIN INVESTMENT BOARD73.6K$1.21M—Held
PROFUND ADVISORS LLC71.6K$1.18M—Held
Universal- Beteiligungs- und Servicegesellschaft mbH70.8K$1.17M—Held
Ishara Investments LP70K$1.16M—Held
PIER 88 INVESTMENT PARTNERS LLC68.2K$1.13M—Held
1492 Capital Management LLC67.5K$1.11M—Held
Capricorn Fund Managers Ltd62.2K$1.03M—Held
Creative Planning60.8K$1M—Held
FRANKLIN RESOURCES INC60.3K$995.1K—Held
JANUS HENDERSON GROUP PLC60K$991.4K—Held
WELLINGTON MANAGEMENT GROUP LLP54.5K$900.5K—Held
TUDOR INVESTMENT CORP ET AL53.7K$886.3K$106.8KAdded$67.8K
INTECH INVESTMENT MANAGEMENT LLC52.6K$869K—Held
ADAR1 Capital Management, LLC52.1K$860.4K—Held
CLOUGH CAPITAL PARTNERS L P51.8K$855.7K—Held
Pathstone Holdings, LLC51.4K$849K—Held
IRONWOOD INVESTMENT MANAGEMENT LLC50.9K$840.9K—Held
Dana Investment Advisors, Inc.50.6K$835.1K—Held
Knights of Columbus Asset Advisors LLC49K$809.1K—Held
Man Group plc49K$808.9K—Held
Zurcher Kantonalbank (Zurich Cantonalbank)47.5K$783.5K—Held
State of New Jersey Common Pension Fund D47.4K$782.7K—Held
MAI Capital Management46.5K$768.1K—Held
Headlands Technologies LLC46.4K$765.8K—Held
JANE STREET GROUP, LLC42.9K$708.6K—Held
AlphaQuest LLC42.9K$708.4K—Held
Jump Financial, LLC40.6K$670.4K—Held
Savvy Advisors, Inc.39K$644.4K—Held
CAPSTONE INVESTMENT ADVISORS, LLC38.8K$640.8K—Held
ENVESTNET ASSET MANAGEMENT INC38.7K$638.4K—Held
ARIZONA STATE RETIREMENT SYSTEM38.6K$638K—Held
NEW YORK STATE COMMON RETIREMENT FUND38.5K$635.3K—Held
Icahn Capital38.4K$634.6K—Held
Police & Firemen's Retirement System of New Jersey36.6K$604.4K—Held
SUSQUEHANNA INTERNATIONAL GROUP, LLP34.8K$574K—Held
STATE BOARD OF ADMINISTRATION OF FLORIDA RETIREMENT SYSTEM34.2K$564.1K—Held
PSP Research LLC31.8K$524.4K—Held
CITIGROUP INC30.9K$510.9K—Held
ProShare Advisors LLC30.7K$506.6K—Held
KLP KAPITALFORVALTNING AS29.9K$493.6K—Held
Erste Asset Management GmbH26.9K$446K—Held
POLEN CAPITAL MANAGEMENT LLC26.9K$444.1K—Held
Quantinno Capital Management LP26.3K$434.4K—Held
RAYMOND JAMES FINANCIAL INC25.6K$422.5K—Held
China Universal Asset Management Co., Ltd.23.9K$394.6K—Held
VOYA INVESTMENT MANAGEMENT LLC23.8K$393K—Held
TEACHER RETIREMENT SYSTEM OF TEXAS20.5K$339.1K—Held
Bayforest Capital Ltd20.4K$337.3K—Held
MARTINGALE ASSET MANAGEMENT L P18.5K$306K—Held
Graham Capital Management, L.P.18.1K$299.2K—Held
SBI Okasan Asset Management Co.Ltd.18K$298.4K—Held
Breakwater Capital Group16.4K$270.2K—Held
BNP PARIBAS FINANCIAL MARKETS15.5K$255.9K—Held
VICTORY CAPITAL MANAGEMENT INC15.1K$248.7K—Held
Virtus Investment Advisers, LLC14.8K$244.3K—Held
Russell Investments Group, Ltd.13.5K$222.3K—Held
Tower Research Capital LLC (TRC)12.9K$213.7K—Held
WINTON GROUP Ltd12.7K$209.7K—Held
Walleye Capital LLC12.7K$208.9K—Held
Perbak Capital Partners LLP12.6K$208.2K—Held
GUGGENHEIM CAPITAL LLC12.5K$205.8K—Held
HighTower Advisors, LLC11.8K$194.1K—Held
Vanguard Personalized Indexing Management, LLC11.7K$193.8K—Held
ROYAL BANK OF CANADA11.3K$187K—Held
Empowered Funds, LLC11.3K$187K—Held
INTRUST BANK NA11.2K$184.3K—Held
Ameritas Investment Partners, Inc.11.1K$182.7K—Held
Renaissance Technologies10.7K$176.7K—Held
LPL Financial LLC10.5K$173.4K—Held
XPONANCE LLC10.5K$173.3K—Held
PNC Financial Services Group, Inc.8.48K$140K—Held
T. Rowe Price Investment Management, Inc.7.25M$119.8K—Held
Sterling Capital Management LLC7.24K$119.5K—Held
Bellevue Group AG7.14K$117.8K—Held
Mirae Asset Global Investments Co., Ltd.7.13K$117.7K—Held
Quarry LP7K$115.6K—Held
NBC SECURITIES, INC.6.54K$107.9K—Held
Persistent Asset Partners Ltd5.67K$93.6K—New
ASSETMARK, INC5.19K$85.6K—Held
Eurizon Capital SGR S.p.A.5.04K$82.7K—Held
PRICE T ROWE ASSOCIATES INC /MD/3.92M$64.7K—Held
Aster Capital Management (DIFC) Ltd3.45K$57K—Held
Pinnacle Wealth Planning Services, Inc.2.96K$48.8K—Held
GF FUND MANAGEMENT CO. LTD.2.88K$47.6K—Held
State of Wyoming2.69K$44.3K—Held
BARTLETT & CO. WEALTH MANAGEMENT LLC1.5K$24.8K—Held
Versant Capital Management, Inc1.48K$24.4K—Held
NISA INVESTMENT ADVISORS, LLC1.45K$23.9K—Held
Smartleaf Asset Management LLC1.31K$21.5K—Held
COMERICA BANK1.28K$21.2K—Held
Rockefeller Capital Management L.P.1.26K$20.9K—Held
SBI Securities Co., Ltd.1.22K$20.2K—Held
OSAIC HOLDINGS, INC.1.07K$17.7K—Held
Mitsubishi UFJ Asset Management Co., Ltd.1K$16.3K—Held
GAMMA Investing LLC962$15.9K—Held
US BANCORP \DE\834$13.8K—Held
QUADRANT CAPITAL GROUP LLC758$12.5K—Held
Nemes Rush Group LLC700$11.6K—Held
LOS ANGELES CAPITAL MANAGEMENT LLC627$10.3K—Held
HUNTINGTON NATIONAL BANK491$8.11K—Held
Optiver Holding B.V.409$6.75K—Held
NewEdge Advisors, LLC400$6.6K—Held
Parallel Advisors, LLC358$5.91K—Held
BNP PARIBAS ASSET MANAGEMENT Holding S.A.298.8K$4.93K—Held
Private Wealth Management Group, LLC296$4.89K—Held
EASTERN BANK282$4.66K—Held
UMB Bank, n.a.262$4.33K—Held
City State Bank250$4.13K—Held
SIGNATUREFD, LLC230$3.8K—Held
EverSource Wealth Advisors, LLC207$3.42K—Held
Farther Finance Advisors, LLC202$3.33K—Held
Comprehensive Financial Planning, Inc./PA200$3.3K—Held
Arax Advisory Partners198$3.27K—Held
ROTHSCHILD INVESTMENT LLC195$3.22K—Held
Parkside Financial Bank & Trust189$3.12K—Held
TEACHERS RETIREMENT SYSTEM OF THE STATE OF KENTUCKY153.8K$2.54K—Held
SG Americas Securities, LLC149.6K$2.47K—Held
MassMutual Private Wealth & Trust, FSB143$2.36K—Held
ORG Partners LLC130$2.15K—Held
ALGERT GLOBAL LLC127K$2.1K—Held
Root Financial Partners, LLC100$1.65K—Held
Your Advocates Ltd., LLP100$1.65K—Held
Allworth Financial LP90$1.49K$179Added$48
DekaBank Deutsche Girozentrale59.8K$1.35K—Held
Johnson Financial Group, Inc.81$1.34K—Held
HRT FINANCIAL LP78.4K$1.29K—Held
Physician Wealth Advisors, Inc.76$1.25K—Held
TD PRIVATE CLIENT WEALTH LLC71$1.17K—Held
True Wealth Design, LLC56$925—Held
Employees Retirement System of Texas47.2K$779—Held
Handelsbanken Fonder AB43.8K$723—Held
Blue Trust, Inc.40$660—Held
Bell Investment Advisors, Inc38$628—Held
Larson Financial Group LLC38$627—Held
Manchester Capital Management LLC36$595—Held
Cornerstone Financial Management LLC34$561—Held
Westside Investment Management, Inc.33$545—Held
Thrivent Financial for Lutherans32.6K$538—Held
SageView Advisory Group, LLC25$413—Held
Sunbelt Securities, Inc.22$363—Held
TT International Asset Management LTD27K$353—Held
Keating Financial Advisory Services, Inc.20$330—Held
CANADA LIFE ASSURANCE Co13.4K$222—Held
PUBLIC EMPLOYEES RETIREMENT ASSOCIATION OF COLORADO12.2K$202—Held
Atlantic Private Wealth, LLC10$165—Held
IFP Advisors, Inc9$149$18Added$18
HFM Investment Advisors, LLC9$149—Held
JFS WEALTH ADVISORS, LLC8$133—Held
Rialto Wealth Management, LLC8$133—Held
Private Wealth Asset Management, LLC8$132—Held
Fortitude Family Office, LLC7$116—Held
KBC Group NV6.27K$103$9Added$38
AMALGAMATED BANK4.16K$69—Held
SJS Investment Consulting Inc.4$67—Held
Bay Harbor Wealth Management, LLC3$49—Held
CWM, LLC2.61K$43—Held
Dunhill Financial, LLC2$35—Held
Global Retirement Partners, LLC1$17—Held
Covestor Ltd579$10—Held
Daiwa Securities Group Inc.475$8—Held
HANTZ FINANCIAL SERVICES, INC.57$1—Held
Avion Wealth40$0—Held
NOMURA ASSET MANAGEMENT CO LTD0$0—Held

Who left

Positions as of Q4 2025. 13F filings arrive up to 45 days after quarter-end and omit shorts. Values are quarter-end marks, as filed. Estimated mark is the price effect on shares still held — not realized P&L or investment advice.