| JANUS HENDERSON GROUP PLC | 1.32M | $85.9M | — | Held |
| RA CAPITAL MANAGEMENT, L.P. | 667.2K | $43.6M | — | Held |
| ADAGE CAPITAL PARTNERS GP, L.L.C. | 639.5K | $41.8M | — | Held |
| Saturn V Capital Management LP | 325.7K | $21.3M | — | Held |
| Vanguard | 293.8K | $19.2M | — | Held |
| BlackRock, Inc. | 254.9K | $16.6M | — | Held |
| ADAR1 Capital Management, LLC | 233.9K | $15.3M | — | Held |
| WELLINGTON MANAGEMENT GROUP LLP | 226.1K | $14.8M | — | Held |
| Siren, L.L.C. | 212.3K | $13.9M | — | Held |
| DEERFIELD MANAGEMENT COMPANY, L.P. | 170.6K | $11.1M | — | Held |
| TCG Crossover Management, LLC | 141.9K | $9.27M | — | Held |
| Polar Capital Holdings Plc | 105.8K | $6.91M | — | Held |
| GEODE CAPITAL MANAGEMENT, LLC | 99K | $6.47M | — | Held |
| Deep Track Capital, LP | 95.4K | $6.23M | — | Held |
| Vivo Capital, LLC | 90.5K | $5.91M | — | Held |
| State Street | 84K | $5.48M | — | Held |
| MILLENNIUM MANAGEMENT LLC | 76.7K | $5.01M | — | Held |
| Bank of New York Mellon Corp | 64.9K | $4.24M | — | Held |
| Alyeska Investment Group, L.P. | 40K | $2.61M | — | Held |
| Driehaus Capital Management LLC | 33.7K | $2.2M | — | Held |
| NEUBERGER BERMAN GROUP LLC | 32.5K | $2.13M | — | Held |
| Balyasny Asset Management L.P. | 31.9K | $2.08M | — | Held |
| NORTHERN TRUST CORP | 30.5K | $1.99M | — | Held |
| MORGAN STANLEY | 27K | $1.76M | −$257.2K | Added$477.8K |
| BARCLAYS PLC | 20.2K | $1.32M | −$246.9K | Added$90.3K |
| CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 14.3K | $932.8K | — | Held |
| CITIGROUP INC | 13.8K | $903.8K | — | Held |
| Renaissance Technologies | 11.4K | $746.4K | — | Held |
| Informed Momentum Co LLC | 11.4K | $742.7K | — | Held |
| Squarepoint Ops LLC | 10.5K | $687K | — | Held |
| GOLDMAN SACHS GROUP INC | 10.4K | $676.6K | — | Held |
| SEI INVESTMENTS CO | 6.93K | $452.3K | — | Held |
| BANK OF AMERICA CORP /DE/ | 6.36K | $415.5K | −$34.8K | Added$242K |
| MOUNTAIN PACIFIC INVESTMENT ADVISERS LLC | 6.1K | $398.3K | — | Held |
| STATE OF WISCONSIN INVESTMENT BOARD | 5.76K | $375.9K | — | Held |
| GERBER, LLC | 5.6K | $365.7K | — | Held |
| Nuveen, LLC | 5.55K | $362.5K | — | Held |
| WELLS FARGO & COMPANY/MN | 5.13K | $334.7K | — | Held |
| Citadel | 5.07K | $331.3K | — | Held |
| RHUMBLINE ADVISERS | 4.99K | $325.8K | — | Held |
| TWO SIGMA INVESTMENTS, LP | 4.99K | $325.8K | — | Held |
| BankChampaign, National Association | 4.51K | $294.5K | — | Held |
| Engineers Gate Manager LP | 4.34K | $283.3K | — | Held |
| UBS Group AG | 4.25K | $277.5K | — | Held |
| Trexquant Investment LP | 3.78K | $246.7K | — | Held |
| Virtus Investment Advisers, LLC | 3.56K | $232.8K | — | Held |
| DEUTSCHE BANK AG\ | 3.37K | $220.3K | — | Held |
| Invesco Ltd. | 3.35K | $219.1K | — | Held |
| TD Asset Management Inc | 3.08K | $201.4K | — | Held |
| Caitong International Asset Management Co., Ltd | 2.88K | $188K | — | Held |
| MetLife Investment Management, LLC | 2.54K | $165.7K | — | Held |
| JPMORGAN CHASE & CO | 2.11K | $137.7K | — | Held |
| MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd. | 2.09K | $136.5K | — | Held |
| AlphaQuest LLC | 1.98K | $129.5K | — | Held |
| CANADA PENSION PLAN INVESTMENT BOARD | 1.4K | $91.4K | — | Held |
| NEW YORK STATE COMMON RETIREMENT FUND | 1.2K | $78.4K | — | Held |
| BNP PARIBAS FINANCIAL MARKETS | 1.01K | $66.3K | — | Held |
| MEEDER ASSET MANAGEMENT INC | 994 | $64.9K | — | Held |
| Police & Firemen's Retirement System of New Jersey | 964 | $62.9K | — | Held |
| NewEdge Advisors, LLC | 600 | $39.2K | — | Held |
| Legal & General Group Plc | 479 | $31.3K | — | Held |
| Fidelity | 358 | $23.4K | — | Held |
| Tower Research Capital LLC (TRC) | 347 | $22.7K | — | Held |
| Russell Investments Group, Ltd. | 341 | $22.3K | — | Held |
| ROYAL BANK OF CANADA | 326 | $22K | — | Held |
| Ameritas Investment Partners, Inc. | 300 | $19.6K | — | Held |
| CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM | 281 | $18.3K | — | Held |
| Allworth Financial LP | 100 | $6.53K | −$1.64K | Cut−$1.64K |
| Sterling Capital Management LLC | 29 | $1.89K | — | Held |
| PRICE T ROWE ASSOCIATES INC /MD/ | 22.7K | $1.49K | — | Held |
| GAMMA Investing LLC | 14 | $914 | — | Held |
| Virtu Financial LLC | 10K | $654 | — | Held |
| AMALGAMATED BANK | 144 | $9 | — | Held |
| CWM, LLC | 5 | $0 | — | Held |