| Nantahala Capital Management, LLC | 4.96M | $26.2M | — | Held |
| BlackRock, Inc. | 3.16M | $16.7M | — | Held |
| Vanguard | 2.76M | $14.5M | — | Held |
| SUVRETTA CAPITAL MANAGEMENT, LLC | 2.68M | $14.1M | — | Held |
| Cable Car Capital, LP | 2.5M | $13.2M | — | Held |
| AIGH Capital Management LLC | 2M | $10.5M | — | Held |
| Vivo Capital, LLC | 1.84M | $9.71M | — | Held |
| GEODE CAPITAL MANAGEMENT, LLC | 1.14M | $5.99M | — | Held |
| KENNEDY CAPITAL MANAGEMENT LLC | 1.05M | $5.53M | — | Held |
| TANG CAPITAL MANAGEMENT LLC | 1M | $5.27M | — | Held |
| NAN FUNG TRINITY (HK) LTD | 956.8K | $5.04M | — | Held |
| State Street | 898.4K | $4.73M | — | Held |
| Propel Bio Management, LLC | 830.8K | $4.38M | — | Held |
| AWM Investment Company, Inc. | 774.1K | $4.08M | — | Held |
| 683 Capital Management, LLC | 770K | $4.06M | — | Held |
| Stonepine Capital Management, LLC | 709.5K | $3.74M | — | Held |
| SUSQUEHANNA INTERNATIONAL GROUP, LLP | 701.3K | $3.7M | — | Held |
| Diametric Capital, LP | 694.4K | $3.66M | — | Held |
| SILVERARC CAPITAL MANAGEMENT, LLC | 627.7K | $3.31M | — | Held |
| Aberdeen Group plc | 616.4K | $3.25M | — | Held |
| Balyasny Asset Management L.P. | 591.7K | $3.12M | — | Held |
| CITIGROUP INC | 521.1K | $2.75M | — | Held |
| BANK OF AMERICA CORP /DE/ | 507.6K | $2.68M | −$5.08K | Cut−$2.14M |
| GOLDMAN SACHS GROUP INC | 502.2K | $2.65M | — | Held |
| Simplify Asset Management Inc. | 500.3K | $2.64M | — | Held |
| MORGAN STANLEY | 473.7K | $2.5M | −$4.74K | Cut−$712.1K |
| NORTHERN TRUST CORP | 419.3K | $2.21M | — | Held |
| J. Goldman & Co LP | 398.9K | $2.1M | — | Held |
| Jain Global LLC | 366.1K | $1.93M | — | Held |
| MILLENNIUM MANAGEMENT LLC | 343.6K | $1.81M | — | Held |
| GROUP ONE TRADING LLC | 324K | $1.71M | — | Held |
| Knoll Capital Management, LLC | 236K | $1.24M | — | Held |
| Bank of New York Mellon Corp | 194.3K | $1.02M | — | Held |
| Pale Fire Capital SE | 174.7K | $920.7K | — | Held |
| Fidelity | 168.5K | $887.8K | — | Held |
| PANAGORA ASSET MANAGEMENT INC | 166.7K | $878.6K | — | Held |
| UBS Group AG | 160.4K | $845.3K | — | Held |
| LOS ANGELES CAPITAL MANAGEMENT LLC | 155K | $787.3K | — | Held |
| CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 132.9K | $700.4K | — | Held |
| Citadel | 127.6K | $672.5K | — | Held |
| Renaissance Technologies | 116.2K | $612.4K | — | Held |
| Icahn Capital | 114.5K | $603.5K | — | Held |
| PRELUDE CAPITAL MANAGEMENT, LLC | 111.2K | $585.9K | — | Held |
| Palo Alto Investors LP | 108.9K | $573.7K | — | Held |
| D. E. Shaw & Co., Inc. | 106K | $558.4K | — | Held |
| BARCLAYS PLC | 99.3K | $523.4K | −$623 | Added$194.4K |
| SIMPLEX TRADING, LLC | 98.2K | $517.7K | — | Held |
| Jump Financial, LLC | 75.9K | $400.2K | — | Held |
| SEI INVESTMENTS CO | 71.2K | $375K | — | Held |
| LPL Financial LLC | 62.9K | $331.3K | — | Held |
| Rangeley Capital, LLC | 57.6K | $303.6K | — | Held |
| Caption Management, LLC | 57K | $300.6K | — | Held |
| Nuveen, LLC | 56.7K | $298.9K | — | Held |
| RHUMBLINE ADVISERS | 55.9K | $294.7K | — | Held |
| XTX Topco Ltd | 49.3K | $260K | — | Held |
| WELLS FARGO & COMPANY/MN | 47.9K | $252.2K | — | Held |
| DEERFIELD MANAGEMENT COMPANY, L.P. | 40K | $210.8K | — | Held |
| DEUTSCHE BANK AG\ | 39.4K | $207.5K | — | Held |
| AXQ CAPITAL, LP | 33.6K | $177K | — | Held |
| Brooklyn Investment Group | 25.8K | $138.4K | — | Held |
| MetLife Investment Management, LLC | 23.9K | $126.2K | — | Held |
| Invesco Ltd. | 23.1K | $121.5K | — | Held |
| Brevan Howard Capital Management LP | 21.7K | $114.4K | — | Held |
| JPMORGAN CHASE & CO | 21.3K | $112.5K | — | Held |
| INTECH INVESTMENT MANAGEMENT LLC | 21.2K | $112K | — | Held |
| MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd. | 20.2K | $106.7K | — | Held |
| Riverwater Partners LLC | 19.4K | $102.1K | — | Held |
| Cetera Investment Advisers | 18.3K | $96.2K | — | Held |
| ALLIANCEBERNSTEIN L.P. | 17.9K | $94.4K | — | Held |
| SHERBROOKE PARK ADVISERS LLC | 15.6K | $82K | — | Held |
| MANUFACTURERS LIFE INSURANCE COMPANY, THE | 13.7K | $71.9K | — | Held |
| ENVESTNET ASSET MANAGEMENT INC | 13.3K | $70.1K | — | Held |
| Legato Capital Management LLC | 13.2K | $69.5K | — | Held |
| Russell Investments Group, Ltd. | 13.1K | $69K | — | Held |
| PVG ASSET MANAGEMENT CORP | 12.6K | $66.2K | — | New |
| Police & Firemen's Retirement System of New Jersey | 11K | $58K | — | Held |
| Truvestments Capital LLC | 10.3K | $54.3K | — | Held |
| BNP PARIBAS FINANCIAL MARKETS | 9.55K | $50.4K | — | Held |
| MAI Capital Management | 5.62K | $29.6K | — | Held |
| Legal & General Group Plc | 4.6K | $24.3K | — | Held |
| Allworth Financial LP | 4.5K | $23.7K | −$45 | Cut−$193 |
| Ameritas Investment Partners, Inc. | 4.46K | $23.5K | — | Held |
| ROYAL BANK OF CANADA | 4.34K | $22K | — | Held |
| Tower Research Capital LLC (TRC) | 3.66K | $19.3K | — | Held |
| CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM | 2.69K | $14.2K | — | Held |
| Farther Finance Advisors, LLC | 2.12K | $11.2K | — | Held |
| OSAIC HOLDINGS, INC. | 1.9K | $10K | — | Held |
| NEW YORK STATE COMMON RETIREMENT FUND | 1.7K | $9.09K | — | Held |
| SHP Wealth Management | 1.67K | $8.79K | — | Held |
| Rockefeller Capital Management L.P. | 1.25K | $6.59K | — | Held |
| JONES FINANCIAL COMPANIES LLLP | 1K | $5.27K | — | Held |
| Sterling Capital Management LLC | 427 | $2.25K | — | Held |
| Parallel Advisors, LLC | 400 | $2.11K | — | Held |
| Vermillion & White Wealth Management Group, LLC | 370 | $1.95K | — | Held |
| UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC | 134 | $706 | — | Held |
| GAMMA Investing LLC | 90 | $474 | — | Held |
| SBI Securities Co., Ltd. | 78 | $411 | — | Held |
| Darwin Wealth Management, LLC | 26 | $137 | — | Held |
| PRICE T ROWE ASSOCIATES INC /MD/ | 24.1K | $128 | — | Held |
| SG Americas Securities, LLC | 10.7K | $57 | — | Held |
| PNC Financial Services Group, Inc. | 5 | $26 | — | Held |
| AMALGAMATED BANK | 1.37K | $7 | — | Held |
| NewEdge Advisors, LLC | 1 | $2 | — | Held |
| CWM, LLC | 55 | $0 | — | Held |