| Nantahala Capital Management, LLC | 5.82M | $38.7M | — | Held |
| BlackRock, Inc. | 4.1M | $27.2M | — | Held |
| Vanguard | 3.34M | $22.2M | — | Held |
| BRAIDWELL LP | 3.11M | $20.6M | — | Held |
| Frazier Life Sciences Management, L.P. | 2.63M | $17.5M | — | Held |
| MILLENNIUM MANAGEMENT LLC | 2.09M | $13.9M | — | Held |
| Stonepine Capital Management, LLC | 1.63M | $10.9M | — | Held |
| State Street | 1.57M | $10.5M | — | Held |
| D. E. Shaw & Co., Inc. | 1.54M | $10.2M | — | Held |
| GEODE CAPITAL MANAGEMENT, LLC | 1.39M | $9.22M | — | Held |
| Citadel | 1.38M | $9.17M | — | Held |
| TANG CAPITAL MANAGEMENT LLC | 1.25M | $8.28M | — | Held |
| RICE HALL JAMES & ASSOCIATES, LLC | 1.15M | $7.67M | — | Held |
| Soleus Capital Management, L.P. | 1.07M | $7.12M | — | Held |
| SCHRODER INVESTMENT MANAGEMENT GROUP | 986.5K | $6.58M | — | Held |
| ARMISTICE CAPITAL, LLC | 976K | $6.49M | — | Held |
| UBS Group AG | 881.4K | $5.86M | — | Held |
| Renaissance Technologies | 799.9K | $5.32M | — | Held |
| PINNACLE ASSOCIATES LTD | 782.4K | $5.2M | — | Held |
| Buckley Capital Advisors, LLC | 744.2K | $4.95M | — | Held |
| MORGAN STANLEY | 730.7K | $4.86M | $135.5K | Added$3.23M |
| EcoR1 Capital, LLC | 675K | $4.49M | — | Held |
| Soviero Asset Management, LP | 655K | $4.36M | — | Held |
| AWH Capital, L.P. | 625K | $4.16M | — | Held |
| BNP PARIBAS FINANCIAL MARKETS | 565K | $3.76M | — | Held |
| CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 517.1K | $3.44M | — | Held |
| NORTHERN TRUST CORP | 445.2K | $2.96M | — | Held |
| DEUTSCHE BANK AG\ | 436.7K | $2.9M | — | Held |
| JANE STREET GROUP, LLC | 417.3K | $2.77M | — | Held |
| BANK OF AMERICA CORP /DE/ | 358K | $2.38M | $135.7K | Added$747.1K |
| GOLDMAN SACHS GROUP INC | 350.7K | $2.33M | — | Held |
| Eversept Partners, LP | 344.1K | $2.29M | — | Held |
| Campbell & CO Investment Adviser LLC | 340.2K | $2.26M | — | Held |
| Assenagon Asset Management S.A. | 270.2K | $1.8M | — | Held |
| STIFEL FINANCIAL CORP | 263.1K | $1.75M | — | Held |
| CM Management, LLC | 250K | $1.66M | — | Held |
| JPMORGAN CHASE & CO | 225.6K | $1.5M | — | Held |
| National Philanthropic Trust | 200K | $1.33M | $102K | Added$102K |
| WELLINGTON MANAGEMENT GROUP LLP | 195.2K | $1.3M | — | Held |
| Squarepoint Ops LLC | 174.3K | $1.16M | — | Held |
| Potomac Capital Management, Inc. | 141.2K | $938.8K | — | Held |
| Bank of New York Mellon Corp | 137.8K | $916.4K | — | Held |
| Walleye Capital LLC | 133K | $884.3K | — | Held |
| BARCLAYS PLC | 130.7K | $869.4K | $54K | Added$219.2K |
| AMERICAN ASSETS INC | 125K | $831.3K | — | Held |
| TWO SIGMA INVESTMENTS, LP | 102.1K | $678.9K | — | Held |
| BOOTHBAY FUND MANAGEMENT, LLC | 101.8K | $677K | — | Held |
| Knott David M Jr | 99.3K | $660.6K | — | Held |
| Quantinno Capital Management LP | 89K | $591.8K | — | Held |
| Nuveen, LLC | 88.4K | $587.7K | — | Held |
| Seven Fleet Capital Management LP | 82K | $545.3K | — | Held |
| SUSQUEHANNA INTERNATIONAL GROUP, LLP | 80.2K | $533K | — | Held |
| Marex Group plc | 76.8K | $511.1K | — | New |
| WELLS FARGO & COMPANY/MN | 76K | $505.2K | — | Held |
| RHUMBLINE ADVISERS | 70K | $465.8K | — | Held |
| XTX Topco Ltd | 68K | $452.1K | — | Held |
| Russell Investments Group, Ltd. | 65.6K | $436K | — | Held |
| STATE OF WISCONSIN INVESTMENT BOARD | 65.1K | $432.9K | — | Held |
| ALLIANCEBERNSTEIN L.P. | 62.3K | $414.6K | — | Held |
| Brooklyn Investment Group | 61.7K | $411.2K | — | Held |
| LPL Financial LLC | 61K | $405.5K | — | Held |
| AMERIPRISE FINANCIAL INC | 54.3K | $360.9K | — | Held |
| GWN SECURITIES INC. | 48.9K | $325.3K | — | Held |
| SEI INVESTMENTS CO | 45.5K | $302.6K | — | Held |
| Man Group plc | 43.4K | $288.6K | — | Held |
| Integrated Quantitative Investments LLC | 42K | $279.1K | — | Held |
| Pathstone Holdings, LLC | 41.1K | $273K | — | Held |
| PRELUDE CAPITAL MANAGEMENT, LLC | 39.3K | $261.4K | — | Held |
| ADAR1 Capital Management, LLC | 35.9K | $238.5K | — | Held |
| CITIGROUP INC | 35.5K | $235.9K | — | Held |
| Invesco Ltd. | 34.1K | $226.9K | — | Held |
| INTECH INVESTMENT MANAGEMENT LLC | 32.7K | $217.4K | — | Held |
| FOX RUN MANAGEMENT, L.L.C. | 32.7K | $217.2K | — | Held |
| Atom Investors LP | 32.5K | $216.1K | — | Held |
| MetLife Investment Management, LLC | 28.4K | $188.8K | — | Held |
| Aquatic Capital Management LLC | 28.4K | $188.7K | — | Held |
| Versor Investments LP | 26.7K | $177.6K | — | Held |
| ExodusPoint Capital Management, LP | 26.2K | $174.3K | — | Held |
| Alpine Global Management, LLC | 26.1K | $173.5K | — | Held |
| SIMPLEX TRADING, LLC | 24.7K | $164.1K | — | Held |
| MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd. | 24.1K | $160.4K | — | Held |
| Creative Planning | 24.1K | $160K | — | Held |
| Icahn Capital | 23K | $152.8K | — | Held |
| OXFORD ASSET MANAGEMENT LLP | 22.9K | $152.4K | — | New |
| MERCER GLOBAL ADVISORS INC /ADV | 21.7K | $144K | — | Held |
| PROFUND ADVISORS LLC | 20.4K | $135.8K | — | Held |
| TWO SIGMA ADVISERS, LP | 19.4K | $129K | — | Held |
| Trexquant Investment LP | 18.8K | $125.3K | — | Held |
| Rafferty Asset Management, LLC | 18.2K | $120.9K | — | Held |
| Quadrature Capital Ltd | 18.1K | $120.3K | — | Held |
| Pure Financial Advisors, LLC | 17.8K | $118.4K | — | Held |
| ENVESTNET ASSET MANAGEMENT INC | 16.7K | $111.1K | — | Held |
| AlphaQuest LLC | 16.4K | $109.1K | — | Held |
| MANUFACTURERS LIFE INSURANCE COMPANY, THE | 16.2K | $107.9K | — | Held |
| Global Retirement Partners, LLC | 16.2K | $107.7K | — | Held |
| Police & Firemen's Retirement System of New Jersey | 15.8K | $105.3K | — | Held |
| Legato Capital Management LLC | 15.1K | $100.6K | — | Held |
| NEW YORK STATE COMMON RETIREMENT FUND | 14.2K | $94.1K | — | Held |
| Sanctuary Advisors, LLC | 13.9K | $92.7K | — | New |
| Tradition Wealth Management, LLC | 12.5K | $83.1K | — | Held |
| ProShare Advisors LLC | 12.3K | $82K | — | Held |
| OSAIC HOLDINGS, INC. | 12.1K | $80.4K | — | Held |
| STATE BOARD OF ADMINISTRATION OF FLORIDA RETIREMENT SYSTEM | 11.9K | $78.9K | — | Held |
| AIRE ADVISORS, LLC | 11.4K | $76.1K | — | Held |
| TWO SIGMA SECURITIES, LLC | 11.3K | $75.1K | — | Held |
| China Universal Asset Management Co., Ltd. | 10.6K | $70.2K | — | Held |
| Aaron Wealth Advisors LLC | 10.3K | $68.2K | — | Held |
| VOYA INVESTMENT MANAGEMENT LLC | 10K | $66.5K | — | Held |
| Allspring Global Investments Holdings, LLC | 10.3K | $64.1K | — | Held |
| State of Wyoming | 9.5K | $63.2K | — | Held |
| NISA INVESTMENT ADVISORS, LLC | 6.82K | $45.4K | — | Held |
| PRIVATE TRUST CO NA | 6.56K | $43.6K | — | Held |
| Federation des caisses Desjardins du Quebec | 6.2K | $41.2K | — | Held |
| Legal & General Group Plc | 5.45K | $36.3K | — | Held |
| Ameritas Investment Partners, Inc. | 5.42K | $36K | — | Held |
| MEEDER ASSET MANAGEMENT INC | 5.06K | $33.7K | — | Held |
| Larson Financial Group LLC | 4.91K | $32.7K | — | Held |
| IFP Advisors, Inc | 4.73K | $31.4K | $2.41K | Added$2.41K |
| ROYAL BANK OF CANADA | 3.39K | $23K | — | Held |
| Farther Finance Advisors, LLC | 3.25K | $21.6K | — | Held |
| MAI Capital Management | 3.2K | $21.3K | — | Held |
| CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM | 3.2K | $21.3K | — | Held |
| Fidelity | 3.15K | $21K | — | Held |
| Mirae Asset Global Investments Co., Ltd. | 3.15K | $20.9K | — | Held |
| Tower Research Capital LLC (TRC) | 2.73K | $18.2K | — | Held |
| Parallel Advisors, LLC | 2.06K | $13.7K | — | Held |
| POM Investment Strategies, LLC | 2K | $13.3K | — | Held |
| GAMMA Investing LLC | 1.92K | $12.8K | — | Held |
| NATIXIS | 1.44K | $9.56K | — | Held |
| GF FUND MANAGEMENT CO. LTD. | 1.27K | $8.46K | — | Held |
| Plante Moran Financial Advisors, LLC | 1.22K | $8.11K | — | Held |
| IAG Wealth Partners, LLC | 927 | $6.17K | — | Held |
| CIBC Private Wealth Group LLC | 750 | $4.99K | — | Held |
| Versant Capital Management, Inc | 539 | $3.58K | — | Held |
| MassMutual Private Wealth & Trust, FSB | 471 | $3.13K | — | Held |
| SUMMIT SECURITIES GROUP LLC | 455 | $3.03K | — | Held |
| HRT FINANCIAL LP | 428.6K | $2.85K | — | Held |
| PNC Financial Services Group, Inc. | 412 | $2.74K | — | Held |
| Olistico Wealth, LLC | 400 | $2.66K | — | Held |
| JONES FINANCIAL COMPANIES LLLP | 390 | $2.59K | — | Held |
| Sterling Capital Management LLC | 381 | $2.53K | — | Held |
| Smartleaf Asset Management LLC | 348 | $2.32K | — | Held |
| LOS ANGELES CAPITAL MANAGEMENT LLC | 316 | $2.12K | — | Held |
| Arax Advisory Partners | 305 | $2.03K | — | Held |
| HUNTINGTON NATIONAL BANK | 301 | $2K | — | Held |
| Stephens Consulting, LLC | 290 | $1.93K | — | Held |
| Optiver Holding B.V. | 277 | $1.84K | — | Held |
| SBI Securities Co., Ltd. | 262 | $1.74K | — | Held |
| Allworth Financial LP | 238 | $1.58K | — | New |
| Rockefeller Capital Management L.P. | 124 | $825 | — | Held |
| GSA CAPITAL PARTNERS LLP | 118.1K | $786 | — | Held |
| TD PRIVATE CLIENT WEALTH LLC | 101 | $672 | — | Held |
| Krensavage Asset Management, LLC | 100K | $665 | — | Held |
| Newbridge Financial Services Group, Inc. | 100 | $665 | — | Held |
| PRICE T ROWE ASSOCIATES INC /MD/ | 85.4K | $568 | — | Held |
| US BANCORP \DE\ | 84 | $559 | — | Held |
| GHP Investment Advisors, Inc. | 80 | $532 | — | Held |
| Caitong International Asset Management Co., Ltd | 37 | $246 | — | Held |
| SIGNATUREFD, LLC | 37 | $246 | — | Held |
| Clearstead Advisors, LLC | 34 | $228 | — | Held |
| SG Americas Securities, LLC | 27.8K | $185 | — | Held |
| Virtu Financial LLC | 22.7K | $151 | — | Held |
| ROTHSCHILD INVESTMENT LLC | 20 | $133 | — | Held |
| CoreCap Advisors, LLC | 19 | $126 | — | Held |
| State of Alaska, Department of Revenue | 4.33K | $28 | — | Held |
| AMALGAMATED BANK | 1.63K | $11 | — | Held |
| Daiwa Securities Group Inc. | 262 | $2 | — | Held |
| CWM, LLC | 112 | $1 | — | Held |
| Covestor Ltd | 6 | $0 | — | Held |