| Vanguard | 5.59M | $152.3M | — | Held |
| BlackRock, Inc. | 5.24M | $142.5M | — | Held |
| FARALLON CAPITAL MANAGEMENT LLC | 5.16M | $140.4M | −$66.7M | Cut−$91.6M |
| State Street | 3.11M | $84.6M | — | Held |
| ARMISTICE CAPITAL, LLC | 3.04M | $82.6M | — | Held |
| Erste Asset Management GmbH | 2.8M | $76.3M | — | Held |
| D. E. Shaw & Co., Inc. | 1.84M | $50M | — | Held |
| NOMURA ASSET MANAGEMENT INTERNATIONAL INC. | 1.7M | $46.3M | — | Held |
| GEODE CAPITAL MANAGEMENT, LLC | 1.41M | $38.5M | — | Held |
| Citadel | 1.35M | $36.7M | — | Held |
| MPM BIOIMPACT LLC | 1.3M | $35.5M | — | Held |
| Frazier Life Sciences Management, L.P. | 1.28M | $34.8M | — | Held |
| TCG Crossover Management, LLC | 1.18M | $32.2M | — | Held |
| SUSQUEHANNA INTERNATIONAL GROUP, LLP | 1.14M | $31.1M | — | Held |
| JANE STREET GROUP, LLC | 1.12M | $30.5M | — | Held |
| GOLDMAN SACHS GROUP INC | 1.03M | $28.1M | — | Held |
| DIMENSIONAL FUND ADVISORS LP | 1M | $27.3M | — | Held |
| MORGAN STANLEY | 894K | $24.3M | −$5.39M | Added$7.6M |
| Rock Springs Capital Management LP | 799.2K | $21.8M | — | Held |
| Capital International Investors | 703.9K | $19.2M | — | Held |
| Balyasny Asset Management L.P. | 635.1K | $17.3M | — | Held |
| SCS Capital Management LLC | 598.8K | $16.3M | — | Held |
| WELLINGTON MANAGEMENT GROUP LLP | 597.8K | $16.3M | — | Held |
| Connor, Clark & Lunn Investment Management Ltd. | 562.6K | $15.3M | — | Held |
| BANK OF AMERICA CORP /DE/ | 539.3K | $14.7M | −$6.97M | Cut−$30.7M |
| Foresite Capital Management VI LLC | 530K | $14.4M | — | Held |
| NORTHERN TRUST CORP | 511.9K | $13.9M | — | Held |
| Artisan Partners Limited Partnership | 504.5K | $13.7M | — | Held |
| CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 482.6K | $13.1M | — | Held |
| ArrowMark Colorado Holdings LLC | 466.2K | $12.7M | — | Held |
| MILLENNIUM MANAGEMENT LLC | 398.3K | $10.8M | — | Held |
| Qube Research & Technologies Ltd | 378.4K | $10.3M | — | Held |
| MARSHALL WACE, LLP | 353.6K | $9.63M | — | Held |
| Legal & General Group Plc | 328.2K | $8.93M | — | Held |
| Vestal Point Capital, LP | 325K | $8.85M | — | Held |
| BARCLAYS PLC | 320.4K | $8.72M | −$2.79M | Added$48.1K |
| TWO SIGMA INVESTMENTS, LP | 300.2K | $8.17M | — | Held |
| Nuveen, LLC | 297.9K | $8.11M | — | Held |
| JPMORGAN CHASE & CO | 251.8K | $6.85M | — | Held |
| BNP PARIBAS FINANCIAL MARKETS | 242.7K | $6.61M | — | Held |
| Trexquant Investment LP | 226.6K | $6.17M | — | Held |
| Palo Alto Investors LP | 212.5K | $5.78M | — | Held |
| Pictet Asset Management Holding SA | 205K | $5.57M | — | Held |
| Point72 Asset Management, L.P. | 200K | $5.44M | — | Held |
| Bank of New York Mellon Corp | 196.9K | $5.36M | — | Held |
| ADAR1 Capital Management, LLC | 188K | $5.12M | — | Held |
| Edgestream Partners, L.P. | 168K | $4.57M | — | Held |
| GROUP ONE TRADING LLC | 166.6K | $4.54M | — | Held |
| Tetragon Partners GP Ltd | 162K | $4.41M | — | Held |
| AMERICAN CENTURY COMPANIES INC | 151.5K | $4.12M | — | Held |
| MOORE CAPITAL MANAGEMENT, LP | 150K | $4.08M | — | Held |
| Allianz Asset Management GmbH | 147.9K | $4.03M | — | Held |
| HighVista Strategies LLC | 147.7K | $4.02M | — | Held |
| Invesco Ltd. | 142.6K | $3.88M | — | Held |
| Exome Asset Management LLC | 136.9K | $3.73M | — | Held |
| UBS Group AG | 126.7K | $3.45M | — | Held |
| DEUTSCHE BANK AG\ | 122.9K | $3.34M | — | Held |
| Squarepoint Ops LLC | 121.8K | $3.31M | — | Held |
| Ikarian Capital, LLC | 115.5K | $3.14M | — | Held |
| JACOBS LEVY EQUITY MANAGEMENT, INC | 111.2K | $3.03M | — | Held |
| Assenagon Asset Management S.A. | 110.4K | $3.01M | — | Held |
| Swiss National Bank | 108.9K | $2.96M | — | Held |
| Rafferty Asset Management, LLC | 107.5K | $2.93M | — | Held |
| UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC | 107K | $2.91M | — | Held |
| Hudson Bay Capital Management LP | 100K | $2.72M | −$1.29M | Cut−$2.2M |
| Parkman Healthcare Partners LLC | 99.6K | $2.71M | — | Held |
| Caption Management, LLC | 93.5K | $2.54M | — | Held |
| CITIGROUP INC | 88.2K | $2.4M | — | Held |
| TWO SIGMA ADVISERS, LP | 85.2K | $2.32M | — | Held |
| PRUDENTIAL FINANCIAL INC | 81.1K | $2.21M | — | Held |
| PANAGORA ASSET MANAGEMENT INC | 79.8K | $2.17M | — | Held |
| Y-Intercept (Hong Kong) Ltd | 79.8K | $2.17M | — | Held |
| Lynx1 Capital Management LP | 79.6K | $2.17M | — | Held |
| FIRST TRUST ADVISORS LP | 78.7K | $2.14M | — | Held |
| Woodline Partners LP | 74.4K | $2.03M | — | Held |
| RHUMBLINE ADVISERS | 74.2K | $2.02M | — | Held |
| ExodusPoint Capital Management, LP | 71.1K | $1.94M | — | Held |
| MANUFACTURERS LIFE INSURANCE COMPANY, THE | 70.3K | $1.92M | — | Held |
| KENNEDY CAPITAL MANAGEMENT LLC | 65.7K | $1.79M | — | Held |
| WELLS FARGO & COMPANY/MN | 64.8K | $1.76M | — | Held |
| ALLIANCEBERNSTEIN L.P. | 61.5K | $1.67M | — | Held |
| TUDOR INVESTMENT CORP ET AL | 61.3K | $1.67M | — | New |
| Universal- Beteiligungs- und Servicegesellschaft mbH | 60.6K | $1.65M | — | Held |
| TD Asset Management Inc | 57.3K | $1.56M | — | Held |
| CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM | 53.2K | $1.45M | — | Held |
| Headlands Technologies LLC | 52.7K | $1.44M | — | Held |
| NEW YORK STATE COMMON RETIREMENT FUND | 51.7K | $1.41M | — | Held |
| Engineers Gate Manager LP | 49.1K | $1.34M | — | Held |
| ALPS ADVISORS INC | 47.9K | $1.3M | — | Held |
| State of New Jersey Common Pension Fund D | 47.4K | $1.29M | — | Held |
| SCHRODER INVESTMENT MANAGEMENT GROUP | 46.2K | $1.26M | — | Held |
| XTX Topco Ltd | 46K | $1.25M | — | Held |
| VIRGINIA RETIREMENT SYSTEMS ET Al | 41.4K | $1.13M | — | Held |
| Freestone Grove Partners LP | 40.8K | $1.11M | — | Held |
| PROFUND ADVISORS LLC | 39.9K | $1.08M | — | Held |
| MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd. | 39.5K | $1.07M | — | Held |
| BOOTHBAY FUND MANAGEMENT, LLC | 37.3K | $1.02M | — | Held |
| Aquatic Capital Management LLC | 36K | $980.1K | — | Held |
| Quadrature Capital Ltd | 35.9K | $976K | — | Held |
| CAPITAL FUND MANAGEMENT S.A. | 35.8K | $974.1K | — | Held |
| EDGEWOOD MANAGEMENT LLC | 35K | $952.7K | — | Held |
| STATE OF WISCONSIN INVESTMENT BOARD | 34.6K | $942.8K | — | Held |
| PRELUDE CAPITAL MANAGEMENT, LLC | 33.7K | $916.9K | — | Held |
| Hennion & Walsh Asset Management, Inc. | 33.5K | $911.1K | — | Held |
| Jump Financial, LLC | 33.1K | $899.7K | — | Held |
| MetLife Investment Management, LLC | 32.2K | $876.9K | — | Held |
| Seven Fleet Capital Management LP | 31.7K | $863.9K | — | Held |
| Ishara Investments LP | 30K | $816.6K | — | Held |
| INTECH INVESTMENT MANAGEMENT LLC | 27.8K | $757.5K | — | Held |
| Icahn Capital | 27.9K | $754K | — | Held |
| VICTORY CAPITAL MANAGEMENT INC | 27.7K | $753.9K | — | Held |
| Quantinno Capital Management LP | 26.9K | $731K | — | Held |
| Aigen Investment Management, LP | 25.9K | $705.4K | — | Held |
| Virtus Investment Advisers, LLC | 25.1K | $682.1K | — | Held |
| FEDERATED HERMES, INC. | 25.1K | $682K | — | Held |
| Russell Investments Group, Ltd. | 24.7K | $671.8K | — | Held |
| Fidelity | 24.3K | $662.4K | — | Held |
| JANUS HENDERSON GROUP PLC | 23.8K | $648.2K | — | Held |
| Zurcher Kantonalbank (Zurich Cantonalbank) | 23.6K | $642.9K | — | Held |
| Rockefeller Capital Management L.P. | 21.1K | $573.6K | — | Held |
| OAK RIDGE INVESTMENTS LLC | 20K | $544.5K | — | Held |
| Brevan Howard Capital Management LP | 19.3K | $526.3K | — | Held |
| Entropy Technologies, LP | 19.2K | $521.6K | — | Held |
| VARCOV Co. | 19.1K | $520K | — | Held |
| CANADA PENSION PLAN INVESTMENT BOARD | 18.2K | $495.4K | — | Held |
| Knott David M Jr | 18K | $490K | — | Held |
| Inspire Investing, LLC | 17.1K | $465.9K | — | Held |
| WINTON GROUP Ltd | 16.8K | $456K | — | Held |
| STATE BOARD OF ADMINISTRATION OF FLORIDA RETIREMENT SYSTEM | 15.8K | $431.4K | — | Held |
| Police & Firemen's Retirement System of New Jersey | 15.6K | $425.7K | — | Held |
| ARIZONA STATE RETIREMENT SYSTEM | 15.3K | $417.3K | — | Held |
| SEI INVESTMENTS CO | 14.4K | $390.7K | — | Held |
| Tidal Investments LLC | 14.2K | $387.8K | — | Held |
| Atria Investments, Inc | 14.1K | $382.5K | — | Held |
| AlphaCentric Advisors LLC | 13.1K | $357.7K | — | Held |
| ROYAL BANK OF CANADA | 13.1K | $354K | — | Held |
| ProShare Advisors LLC | 12.9K | $351.1K | — | Held |
| ENVESTNET ASSET MANAGEMENT INC | 12.7K | $346.6K | — | Held |
| COMMERCE BANK | 12.7K | $344.5K | — | Held |
| Bridgefront Capital, LLC | 12.6K | $344.3K | — | Held |
| FOX RUN MANAGEMENT, L.L.C. | 12.6K | $342.1K | — | Held |
| NOMURA HOLDINGS INC | 12.4K | $336.3K | — | Held |
| AlphaQuest LLC | 12K | $325.6K | — | Held |
| Pale Fire Capital SE | 11.8K | $322.6K | — | Held |
| KLP KAPITALFORVALTNING AS | 11.8K | $321.2K | — | Held |
| Gallagher Fiduciary Advisors, LLC | 11.2K | $306.1K | — | Held |
| Creative Planning | 10.4K | $282.3K | — | Held |
| CAPSTONE INVESTMENT ADVISORS, LLC | 10.4K | $281.9K | — | Held |
| VOYA INVESTMENT MANAGEMENT LLC | 10.1K | $275.8K | — | Held |
| STRS OHIO | 10K | $272.2K | — | Held |
| China Universal Asset Management Co., Ltd. | 9.5K | $258.6K | — | Held |
| TWO SIGMA SECURITIES, LLC | 9.2K | $250.4K | — | Held |
| DIVERSIFIED TRUST CO | 9.16K | $249.4K | — | Held |
| Inscription Capital, LLC | 8.6K | $234K | — | Held |
| Fiduciary Trust Co | 8.25K | $224.5K | — | Held |
| Campbell & CO Investment Adviser LLC | 8.23K | $223.9K | — | Held |
| TEACHER RETIREMENT SYSTEM OF TEXAS | 8.17K | $222.3K | — | Held |
| GRIFFIN ASSET MANAGEMENT, INC. | 7.38K | $200.9K | — | Held |
| Caitong International Asset Management Co., Ltd | 6.74K | $183.5K | — | Held |
| Tower Research Capital LLC (TRC) | 6.63K | $180.6K | — | Held |
| Ensign Peak Advisors, Inc | 6.57K | $178.8K | — | Held |
| SIMPLEX TRADING, LLC | 6.21K | $169K | — | Held |
| CONTRAVISORY INVESTMENT MANAGEMENT, INC. | 6.19K | $168.5K | — | Held |
| Ameritas Investment Partners, Inc. | 4.72K | $128.5K | — | Held |
| PNC Financial Services Group, Inc. | 4.71K | $128.2K | — | Held |
| Sterling Capital Management LLC | 3.28K | $89.4K | — | Held |
| FIRST HORIZON CORP | 3.1K | $84.2K | — | Held |
| Quarry LP | 3K | $81.7K | — | Held |
| Mirae Asset Global Investments Co., Ltd. | 2.83K | $77.1K | — | Held |
| Eurizon Capital SGR S.p.A. | 2.65K | $72.3K | — | Held |
| OSAIC HOLDINGS, INC. | 1.75K | $47.6K | — | Held |
| BNP PARIBAS ASSET MANAGEMENT Holding S.A. | 1.36M | $37K | — | Held |
| ASSETMARK, INC | 1.32K | $35.8K | — | Held |
| GW&K Investment Management, LLC | 1.21M | $32.9K | — | Held |
| GF FUND MANAGEMENT CO. LTD. | 1.15K | $31.2K | — | Held |
| PRICE T ROWE ASSOCIATES INC /MD/ | 637.9K | $17.4K | — | Held |
| Torren Management, LLC | 615 | $16.7K | — | Held |
| US BANCORP \DE\ | 509 | $13.9K | — | Held |
| CITY HOLDING CO | 500 | $13.6K | — | Held |
| Hurley Capital, LLC | 486 | $13.2K | — | Held |
| LOS ANGELES CAPITAL MANAGEMENT LLC | 300 | $13K | — | Held |
| Smartleaf Asset Management LLC | 460 | $12.6K | — | Held |
| ALGERT GLOBAL LLC | 414.2K | $11.3K | — | Held |
| SIGNATUREFD, LLC | 399 | $10.9K | — | Held |
| Quantbot Technologies LP | 386 | $10.5K | — | Held |
| Aspect Partners, LLC | 361 | $9.83K | — | Held |
| NISA INVESTMENT ADVISORS, LLC | 295 | $8.03K | — | Held |
| Bell Investment Advisors, Inc | 281 | $7.65K | — | Held |
| THURSTON, SPRINGER, MILLER, HERD & TITAK, INC. | 241 | $6.56K | — | Held |
| COMERICA BANK | 209 | $5.7K | — | Held |
| Hillsdale Investment Management Inc. | 200 | $5.44K | — | Held |
| Concord Wealth Partners | 200 | $5.44K | — | Held |
| GAMMA Investing LLC | 171 | $4.66K | — | Held |
| Farther Finance Advisors, LLC | 151 | $4.11K | — | Held |
| HRT FINANCIAL LP | 146.5K | $3.99K | — | Held |
| BROWN BROTHERS HARRIMAN & CO | 144 | $3.92K | — | Held |
| Triumph Capital Management | 100 | $2.72K | — | New |
| Versant Capital Management, Inc | 92 | $2.5K | — | Held |
| Private Wealth Management Group, LLC | 83 | $2.26K | — | Held |
| Arax Advisory Partners | 79 | $2.15K | — | Held |
| Root Financial Partners, LLC | 66 | $1.8K | — | Held |
| DekaBank Deutsche Girozentrale | 40K | $1.27K | — | Held |
| SG Americas Securities, LLC | 46.5K | $1.26K | — | Held |
| Pinnacle Wealth Planning Services, Inc. | 40 | $1.09K | — | Held |
| HARBOR CAPITAL ADVISORS, INC. | 37.2K | $1.01K | — | Held |
| EverSource Wealth Advisors, LLC | 34 | $926 | — | Held |
| Keating Financial Advisory Services, Inc. | 21 | $572 | — | Held |
| Handelsbanken Fonder AB | 19.3K | $526 | — | Held |
| Manchester Capital Management LLC | 16 | $436 | — | Held |
| QUADRANT CAPITAL GROUP LLC | 16 | $436 | — | Held |
| Parkside Financial Bank & Trust | 15 | $409 | — | Held |
| Parallel Advisors, LLC | 12 | $327 | — | Held |
| Allworth Financial LP | 11 | $300 | −$142 | Cut−$383 |
| Global Retirement Partners, LLC | 11 | $299 | — | Held |
| CANADA LIFE ASSURANCE Co | 10.4K | $284 | — | Held |
| SageView Advisory Group, LLC | 10 | $273 | — | Held |
| Steward Partners Investment Advisory, LLC | 9 | $245 | — | Held |
| PUBLIC EMPLOYEES RETIREMENT ASSOCIATION OF COLORADO | 6.8K | $185 | — | Held |
| IFP Advisors, Inc | 6 | $163 | −$78 | Cut−$78 |
| Fortitude Family Office, LLC | 5 | $136 | — | Held |
| ROTHSCHILD INVESTMENT LLC | 5 | $136 | — | Held |
| Essential Partners LLC | 5 | $136 | −$65 | Cut−$4.48K |
| State of Alaska, Department of Revenue | 5K | $136 | — | Held |
| JFS WEALTH ADVISORS, LLC | 4 | $109 | — | Held |
| SBI Securities Co., Ltd. | 4 | $109 | — | Held |
| Bay Harbor Wealth Management, LLC | 4 | $108 | — | Held |
| KBC Group NV | 3.02K | $82 | −$22 | Added$12 |
| Rialto Wealth Management, LLC | 2 | $55 | — | Held |
| Dunhill Financial, LLC | 2 | $50 | — | Held |
| AMALGAMATED BANK | 1.77K | $48 | — | Held |
| SJS Investment Consulting Inc. | 1 | $28 | — | Held |
| JONES FINANCIAL COMPANIES LLLP | 1 | $27 | — | Held |
| T. Rowe Price Investment Management, Inc. | 892 | $25 | — | Held |
| CWM, LLC | 484 | $13 | — | Held |
| Daiwa Securities Group Inc. | 204 | $6 | — | Held |
| BESSEMER GROUP INC | 76 | $2 | — | Held |
| Covestor Ltd | 41 | $1 | — | Held |