| BAKER BROS. ADVISORS LP | 5.07M | $52.1M | — | Held |
| 5AM Venture Management, LLC | 4.06M | $41.7M | — | Held |
| BlackRock, Inc. | 2.15M | $22.1M | — | Held |
| TCG Crossover Management, LLC | 2.09M | $21.5M | — | Held |
| Merck & Co., Inc. | 1.74M | $17.9M | — | Held |
| Vanguard | 1.63M | $16.8M | — | Held |
| WELLINGTON MANAGEMENT GROUP LLP | 1.04M | $10.7M | — | Held |
| StepStone Group LP | 761.3K | $7.83M | — | Held |
| GEODE CAPITAL MANAGEMENT, LLC | 639.9K | $6.58M | — | Held |
| State Street | 508.8K | $5.23M | — | Held |
| MILLENNIUM MANAGEMENT LLC | 497.7K | $5.12M | — | Held |
| JACOBS LEVY EQUITY MANAGEMENT, INC | 362.8K | $3.73M | — | Held |
| Citadel | 292.8K | $3.01M | — | Held |
| DIMENSIONAL FUND ADVISORS LP | 273.4K | $2.81M | — | Held |
| MORGAN STANLEY | 260.3K | $2.68M | $1.17M | Added$664.2K |
| Renaissance Technologies | 252.6K | $2.6M | — | Held |
| ROYCE & ASSOCIATES LP | 247.1K | $2.54M | — | Held |
| NORTHERN TRUST CORP | 226.3K | $2.33M | — | Held |
| CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 221.3K | $2.27M | — | Held |
| GOLDMAN SACHS GROUP INC | 205.2K | $2.11M | — | Held |
| BNP PARIBAS FINANCIAL MARKETS | 181K | $1.86M | — | Held |
| Qube Research & Technologies Ltd | 176K | $1.81M | — | Held |
| Woodline Partners LP | 142.1K | $1.46M | — | Held |
| Trexquant Investment LP | 121.3K | $1.25M | — | Held |
| XTX Topco Ltd | 117.7K | $1.21M | — | Held |
| SPHERA FUNDS MANAGEMENT LTD. | 110.9K | $1.14M | — | Held |
| MOORE CAPITAL MANAGEMENT, LP | 105K | $1.08M | — | Held |
| JANE STREET GROUP, LLC | 85.7K | $880.6K | — | Held |
| AMERIPRISE FINANCIAL INC | 84.9K | $872.7K | — | Held |
| Bank of New York Mellon Corp | 79.4K | $816K | — | Held |
| AMERICAN CENTURY COMPANIES INC | 75.3K | $773.9K | — | Held |
| JPMORGAN CHASE & CO | 70.8K | $727.4K | — | Held |
| Quantinno Capital Management LP | 67.7K | $696.1K | — | Held |
| BARCLAYS PLC | 67.1K | $690.3K | $271.6K | Added$338.6K |
| D. E. Shaw & Co., Inc. | 66.9K | $687.5K | — | Held |
| BANK OF AMERICA CORP /DE/ | 66K | $678.3K | $295.6K | Cut−$55.1K |
| CANTOR FITZGERALD, L. P. | 57K | $586K | −$2.72M | Cut−$3.32M |
| Nuveen, LLC | 56.4K | $580K | — | Held |
| MANUFACTURERS LIFE INSURANCE COMPANY, THE | 56.5K | $572.3K | — | Held |
| Lynx1 Capital Management LP | 51.7K | $531.4K | — | Held |
| TWO SIGMA INVESTMENTS, LP | 49.3K | $506.8K | — | Held |
| Nan Fung Group Holdings Ltd | 49.1K | $504.6K | — | Held |
| Brevan Howard Capital Management LP | 48K | $493.1K | — | Held |
| GREAT LAKES ADVISORS, LLC | 43.3K | $444.8K | — | Held |
| Invesco Ltd. | 41K | $421.8K | — | Held |
| BRIDGEWAY CAPITAL MANAGEMENT, LLC | 40K | $411.2K | — | Held |
| TD Asset Management Inc | 37.8K | $388.5K | — | Held |
| PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO | 37.5K | $385.5K | — | Held |
| ExodusPoint Capital Management, LP | 37.4K | $384.8K | — | Held |
| MARTINGALE ASSET MANAGEMENT L P | 37.1K | $380.9K | — | Held |
| RHUMBLINE ADVISERS | 36.2K | $372.4K | — | Held |
| WELLS FARGO & COMPANY/MN | 32.2K | $330.6K | — | Held |
| LOS ANGELES CAPITAL MANAGEMENT LLC | 46K | $317.1K | — | Held |
| Icahn Capital | 26.3K | $269.9K | — | Held |
| CANADA PENSION PLAN INVESTMENT BOARD | 25.6K | $263.2K | — | Held |
| Dynamic Technology Lab Private Ltd | 25.3K | $260K | — | Held |
| ALLIANCEBERNSTEIN L.P. | 24.3K | $250K | — | Held |
| Creative Planning | 22.1K | $227K | — | Held |
| Point72 Asset Management, L.P. | 21.9K | $224.8K | — | Held |
| HSBC HOLDINGS PLC | 20.8K | $216.2K | — | Held |
| DEUTSCHE BANK AG\ | 20.8K | $214K | — | Held |
| MARSHALL WACE, LLP | 20.7K | $212.4K | — | Held |
| Quadrature Capital Ltd | 20.3K | $208.9K | — | Held |
| UBS Group AG | 19K | $195.7K | — | Held |
| MetLife Investment Management, LLC | 17.8K | $183K | — | Held |
| Allspring Global Investments Holdings, LLC | 17.5K | $175.8K | — | Held |
| Fidelity | 16.8K | $172.9K | — | Held |
| AlphaQuest LLC | 15.6K | $160.7K | — | Held |
| SUSQUEHANNA INTERNATIONAL GROUP, LLP | 15.4K | $157.9K | — | Held |
| Balyasny Asset Management L.P. | 14K | $144.1K | — | Held |
| Walleye Capital LLC | 13.5K | $139.1K | — | Held |
| Magnetar Financial LLC | 13.4K | $137.5K | — | Held |
| MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd. | 13.2K | $136.2K | — | Held |
| INTECH INVESTMENT MANAGEMENT LLC | 11.9K | $122.6K | — | Held |
| Inscription Capital, LLC | 11.8K | $121.7K | — | Held |
| Russell Investments Group, Ltd. | 11.6K | $118.9K | — | Held |
| IHT Wealth Management, LLC | 11.1K | $114.1K | — | Held |
| AXQ CAPITAL, LP | 10.7K | $109.5K | — | Held |
| Police & Firemen's Retirement System of New Jersey | 9.47K | $97.4K | — | Held |
| NEW YORK STATE COMMON RETIREMENT FUND | 8.88K | $91.3K | — | Held |
| China Universal Asset Management Co., Ltd. | 6.23K | $64K | — | Held |
| CITIGROUP INC | 4.12K | $42.4K | — | Held |
| Versant Capital Management, Inc | 4.05K | $41.6K | — | Held |
| Legal & General Group Plc | 2.56K | $26.3K | — | Held |
| Ameritas Investment Partners, Inc. | 2.42K | $24.8K | — | Held |
| Tower Research Capital LLC (TRC) | 1.94K | $19.9K | — | Held |
| Mirae Asset Global Investments Co., Ltd. | 1.86K | $19.1K | — | Held |
| CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM | 1.75K | $18K | — | Held |
| Sterling Capital Management LLC | 1.64K | $16.9K | — | Held |
| PNC Financial Services Group, Inc. | 1.14K | $11.7K | — | Held |
| ROYAL BANK OF CANADA | 987 | $11K | — | Held |
| Arax Advisory Partners | 918 | $9.44K | — | Held |
| GF FUND MANAGEMENT CO. LTD. | 751 | $7.72K | — | Held |
| Zurcher Kantonalbank (Zurich Cantonalbank) | 432 | $4.44K | — | Held |
| EverSource Wealth Advisors, LLC | 376 | $3.87K | — | Held |
| Optiver Holding B.V. | 346 | $3.56K | — | Held |
| PRICE T ROWE ASSOCIATES INC /MD/ | 327.4K | $3.37K | — | Held |
| ACADIAN ASSET MANAGEMENT LLC | 310.8K | $3.19K | — | Held |
| Private Wealth Management Group, LLC | 212 | $2.18K | — | Held |
| OSAIC HOLDINGS, INC. | 195 | $2K | — | Held |
| MassMutual Private Wealth & Trust, FSB | 186 | $1.91K | — | Held |
| SIGNATUREFD, LLC | 167 | $1.72K | — | Held |
| NISA INVESTMENT ADVISORS, LLC | 161 | $1.66K | — | Held |
| ALGERT GLOBAL LLC | 128.6K | $1.32K | — | Held |
| GAMMA Investing LLC | 95 | $977 | — | Held |
| Cornerstone Financial Management LLC | 91 | $935 | — | Held |
| Essential Partners LLC | 51 | $524 | $49 | Added$460 |
| UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC | 22 | $226 | — | Held |
| CWM, LLC | 2.11K | $22 | — | Held |
| ROTHSCHILD INVESTMENT LLC | 2 | $21 | — | Held |
| True Wealth Design, LLC | 1 | $11 | — | Held |
| AMALGAMATED BANK | 894 | $9 | — | Held |
| Daiwa Securities Group Inc. | 150 | $2 | — | Held |
| Covestor Ltd | 99 | $1 | — | Held |