| Lynx1 Capital Management LP | 8.02M | $8.02M | — | Held |
| BVF INC/IL | 5.23M | $5.23M | — | Held |
| BAKER BROS. ADVISORS LP | 2.78M | $2.78M | — | Held |
| Vanguard | 2.48M | $2.48M | — | Held |
| BlackRock, Inc. | 1.85M | $1.85M | — | Held |
| Aberdeen Group plc | 1.79M | $1.79M | — | Held |
| Deer Management Co. LLC | 1.25M | $1.25M | — | Held |
| Renaissance Technologies | 1.14M | $1.14M | — | Held |
| TWO SIGMA INVESTMENTS, LP | 651.2K | $651.2K | — | Held |
| DC Funds, LP | 630K | $630K | — | Held |
| GEODE CAPITAL MANAGEMENT, LLC | 562.2K | $562.4K | — | Held |
| MILLENNIUM MANAGEMENT LLC | 403.6K | $403.6K | — | Held |
| JANE STREET GROUP, LLC | 388.3K | $388.3K | — | Held |
| TWO SIGMA ADVISERS, LP | 353.9K | $353.9K | — | Held |
| MARSHALL WACE, LLP | 339.2K | $339.2K | — | Held |
| MORGAN STANLEY | 225.8K | $225.8K | −$185.2K | Cut−$1.32M |
| XTX Topco Ltd | 219K | $219K | — | Held |
| SUSQUEHANNA INTERNATIONAL GROUP, LLP | 216.1K | $216.1K | — | Held |
| State Street | 207.3K | $207.3K | — | Held |
| JACOBS LEVY EQUITY MANAGEMENT, INC | 188.2K | $188.2K | — | Held |
| Citadel | 162.9K | $162.9K | — | Held |
| BROWN BROTHERS HARRIMAN & CO | 143.1K | $143.1K | — | Held |
| BRIDGEWAY CAPITAL MANAGEMENT, LLC | 115.4K | $115.4K | — | Held |
| NORTHERN TRUST CORP | 98.2K | $98.2K | — | Held |
| LETKO, BROSSEAU & ASSOCIATES INC | 95.9K | $95.9K | — | Held |
| NewEdge Wealth, LLC | 91.5K | $91.5K | — | Held |
| GOLDMAN SACHS GROUP INC | 86.9K | $86.9K | — | Held |
| DIMENSIONAL FUND ADVISORS LP | 74.5K | $74.5K | — | Held |
| STIFEL FINANCIAL CORP | 72.4K | $72.4K | — | Held |
| D. E. Shaw & Co., Inc. | 69.4K | $69.4K | — | Held |
| UBS Group AG | 55.2K | $55.2K | — | Held |
| TANG CAPITAL MANAGEMENT LLC | 46.1K | $46.1K | — | Held |
| BANK OF AMERICA CORP /DE/ | 38.7K | $38.7K | −$31.7K | Cut−$367.2K |
| DC Investments Management, LLC | 35.6K | $35.6K | — | Held |
| JPMORGAN CHASE & CO | 33.3K | $33.3K | — | Held |
| BNP PARIBAS FINANCIAL MARKETS | 26.4K | $26.4K | — | Held |
| PANAGORA ASSET MANAGEMENT INC | 25.3K | $25.3K | — | Held |
| HSBC HOLDINGS PLC | 24.6K | $24.8K | — | Held |
| PRUDENTIAL FINANCIAL INC | 24.4K | $24.4K | — | Held |
| BARCLAYS PLC | 23.2K | $23.2K | −$15.6K | Cut−$11.5K |
| LPL Financial LLC | 22.5K | $22.5K | — | Held |
| NEUBERGER BERMAN GROUP LLC | 18.4K | $18.4K | — | Held |
| R Squared Ltd | 16K | $16K | — | Held |
| TWO SIGMA SECURITIES, LLC | 15.9K | $15.9K | — | Held |
| Vanguard Personalized Indexing Management, LLC | 14.7K | $14.7K | — | Held |
| Point72 Asset Management, L.P. | 14.5K | $14.5K | — | Held |
| OSAIC HOLDINGS, INC. | 12.2K | $12.2K | — | Held |
| MIZUHO SECURITIES USA LLC | 10K | $10.1K | — | Held |
| Waldron Private Wealth LLC | 10K | $10K | — | Held |
| Aries Wealth Management | 10K | $10K | — | Held |
| Tower Research Capital LLC (TRC) | 6.77K | $6.77K | — | Held |
| FEDERATED HERMES, INC. | 4.99K | $4.99K | — | Held |
| Fidelity | 500 | $500 | — | Held |
| SBI Securities Co., Ltd. | 392 | $392 | — | Held |
| Strategic Wealth Partners, Ltd. | 311 | $311 | — | Held |
| GSA CAPITAL PARTNERS LLP | 226.2K | $226 | — | Held |
| Rockefeller Capital Management L.P. | 200 | $200 | — | Held |
| ACADIAN ASSET MANAGEMENT LLC | 29.5K | $28 | — | Held |
| HRT FINANCIAL LP | 26.6K | $26 | — | Held |
| NORTHWESTERN MUTUAL WEALTH MANAGEMENT CO | 8 | $8 | — | Held |
| ROYAL BANK OF CANADA | 374 | $0 | — | Held |