| Granahan Investment Management, LLC | 11.6M | $105.8M | — | Held |
| BlackRock, Inc. | 7.55M | $68.9M | — | Held |
| Vanguard | 7.31M | $66.8M | — | Held |
| SUSQUEHANNA ADVISORS GROUP, INC. | 45.4M | $51.1M | — | Held |
| Park West Asset Management LLC | 5.05M | $46.1M | — | Held |
| Context Capital Management, LLC | 36.5M | $41.2M | — | Held |
| GILDER GAGNON HOWE & CO LLC | 4.35M | $39.7M | — | Held |
| D. E. Shaw & Co., Inc. | 2.77M | $25.3M | — | Held |
| Invesco Ltd. | 2.48M | $22.6M | — | Held |
| Driehaus Capital Management LLC | 2.47M | $22.5M | — | Held |
| DeepCurrents Investment Group LLC | 19.1M | $21.5M | — | Held |
| GEODE CAPITAL MANAGEMENT, LLC | 2.29M | $21M | — | Held |
| State Street | 2.22M | $20.3M | — | Held |
| SILVERBACK ASSET MANAGEMENT LLC | 17.5M | $19.9M | — | Held |
| Portolan Capital Management, LLC | 2.08M | $19M | — | Held |
| Divisadero Street Capital Management, LP | 1.62M | $14.8M | — | Held |
| HENNESSY ADVISORS INC | 1.45M | $13.2M | — | Held |
| Zweig-DiMenna Associates LLC | 1.33M | $12.1M | — | Held |
| WELLINGTON MANAGEMENT GROUP LLP | 1.25M | $11.4M | — | Held |
| DIMENSIONAL FUND ADVISORS LP | 1.08M | $9.9M | — | Held |
| TWO SIGMA INVESTMENTS, LP | 1.06M | $9.72M | — | Held |
| MORGAN STANLEY | 947.6K | $8.65M | −$7.25M | Cut−$10.2M |
| G2 Investment Partners Management LLC | 899K | $8.21M | — | Held |
| JPMORGAN CHASE & CO | 865.8K | $7.9M | — | Held |
| NORTHERN TRUST CORP | 847K | $7.73M | — | Held |
| GOLDMAN SACHS GROUP INC | 2.6M | $7.69M | — | Held |
| CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 796.6K | $7.27M | — | Held |
| WELLS FARGO & COMPANY/MN | 607.6K | $5.55M | — | Held |
| MOODY NATIONAL BANK TRUST DIVISION | 593.8K | $5.42M | — | Held |
| UBS Group AG | 536K | $4.89M | — | Held |
| Beartown Capital Management, LLC | 450K | $4.11M | — | Held |
| AYAL Capital Advisors Ltd | 406.5K | $3.71M | — | Held |
| ASHFORD CAPITAL MANAGEMENT INC | 404.7K | $3.69M | — | Held |
| ADVENT CAPITAL MANAGEMENT /DE/ | 3.15M | $3.6M | — | Held |
| CAPSTONE INVESTMENT ADVISORS, LLC | 3M | $3.39M | — | Held |
| UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC | 371.1K | $3.39M | — | Held |
| TUDOR INVESTMENT CORP ET AL | 368K | $3.36M | −$1.95M | Cut−$928K |
| PANAGORA ASSET MANAGEMENT INC | 361.1K | $3.3M | — | Held |
| FIRST TRUST ADVISORS LP | 340.5K | $3.11M | — | Held |
| Bank of New York Mellon Corp | 317K | $2.89M | — | Held |
| BANK OF MONTREAL /CAN/ | 315.5K | $2.88M | — | Held |
| Citadel | 311.5K | $2.84M | — | Held |
| Icahn Capital | 309.7K | $2.83M | — | Held |
| Alpha Wave Global, LP | 286.8K | $2.62M | — | Held |
| Trexquant Investment LP | 279.4K | $2.55M | — | Held |
| Schneider Downs Wealth Management Advisors, LP | 274.8K | $2.51M | — | Held |
| BANK OF AMERICA CORP /DE/ | 270K | $2.46M | −$2.07M | Cut−$3.52M |
| Veradace Capital Management LLC | 240.2K | $2.19M | — | Held |
| Fidelity | 234.9K | $2.14M | — | Held |
| BOOTHBAY FUND MANAGEMENT, LLC | 233.2K | $2.13M | — | Held |
| Russell Investments Group, Ltd. | 217.1K | $1.98M | — | Held |
| CITIZENS FINANCIAL GROUP INC/RI | 212.1K | $1.94M | — | Held |
| Viewpoint Capital Management LLC | 208.5K | $1.9M | — | Held |
| Nuveen, LLC | 184.9K | $1.69M | — | Held |
| BARCLAYS PLC | 177.8K | $1.62M | −$1.04M | Cut−$659.9K |
| STATE OF WISCONSIN INVESTMENT BOARD | 174.2K | $1.59M | — | Held |
| Verition Fund Management LLC | 141.8K | $1.29M | — | Held |
| TWO SIGMA ADVISERS, LP | 136.7K | $1.25M | — | Held |
| Swiss National Bank | 133.7K | $1.22M | — | Held |
| JANE STREET GROUP, LLC | 129.3K | $1.18M | — | Held |
| Progeny 3, Inc. | 128.9K | $1.18M | — | Held |
| Balyasny Asset Management L.P. | 126.5K | $1.16M | — | Held |
| AE Wealth Management LLC | 124.6K | $1.14M | — | Held |
| D.A. DAVIDSON & CO. | 121.4K | $1.11M | — | Held |
| SUSQUEHANNA INTERNATIONAL GROUP, LLP | 119.5K | $1.09M | — | Held |
| HEALTHCARE OF ONTARIO PENSION PLAN TRUST FUND | 118K | $1.08M | — | Held |
| LAZARD ASSET MANAGEMENT LLC | 110.3K | $1.01M | — | Held |
| K2 PRINCIPAL FUND, L.P. | 110K | $1M | — | Held |
| Squarepoint Ops LLC | 103K | $940.3K | — | Held |
| RHUMBLINE ADVISERS | 99.6K | $909.3K | — | Held |
| RAM Investment Partners, LLC | 95.5K | $871.8K | — | Held |
| FRANKLIN RESOURCES INC | 92.4K | $844K | — | Held |
| Clarius Group, LLC | 88.9K | $811.4K | — | Held |
| ALLIANCEBERNSTEIN L.P. | 80.4K | $734.1K | — | Held |
| Ares Systematic Credit Ltd | 78.2K | $714.2K | — | Held |
| NATIXIS | 73.1K | $667.4K | — | Held |
| FULLER & THALER ASSET MANAGEMENT, INC. | 69.3K | $632.8K | — | Held |
| ROYAL BANK OF CANADA | 67.4K | $615K | — | Held |
| MANUFACTURERS LIFE INSURANCE COMPANY, THE | 66.4K | $606.3K | — | Held |
| Centiva Capital, LP | 65K | $593.5K | — | Held |
| CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM | 64.1K | $585.3K | — | Held |
| DEUTSCHE BANK AG\ | 61.7K | $563.3K | — | Held |
| Round Rock Advisors LLC | 60.6K | $553.3K | — | Held |
| Legal & General Group Plc | 59.9K | $547K | — | Held |
| Voleon Capital Management LP | 56.6K | $516.9K | — | Held |
| POLEN CAPITAL MANAGEMENT LLC | 53.7K | $490.1K | — | Held |
| Werba Rubin Papier Wealth Management | 53.3K | $486.5K | — | Held |
| Freestone Grove Partners LP | 53K | $483.8K | — | Held |
| Quantbot Technologies LP | 50.1K | $457.5K | — | Held |
| Numerai GP LLC | 50K | $456.3K | — | Held |
| IRONWOOD INVESTMENT MANAGEMENT LLC | 49.7K | $453.5K | — | Held |
| TEACHER RETIREMENT SYSTEM OF TEXAS | 49.6K | $452.7K | — | Held |
| MetLife Investment Management, LLC | 48.9K | $446.4K | — | Held |
| PRUDENTIAL FINANCIAL INC | 48.2K | $440.2K | — | Held |
| CANADA PENSION PLAN INVESTMENT BOARD | 46.7K | $426.4K | — | Held |
| GROUP ONE TRADING LLC | 41.4K | $378.4K | — | Held |
| LPL Financial LLC | 40.7K | $371.5K | — | Held |
| MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd. | 40.3K | $367.5K | — | Held |
| Parallax Volatility Advisers, L.P. | 40K | $365.2K | — | Held |
| EMERALD MUTUAL FUND ADVISERS TRUST | 39.6K | $361.3K | — | Held |
| PharVision Advisers, LLC | 39.1K | $356.8K | — | Held |
| AMERIPRISE FINANCIAL INC | 38.9K | $355.5K | — | Held |
| Aquatic Capital Management LLC | 38.7K | $353.2K | — | Held |
| SEI INVESTMENTS CO | 34.5K | $315.3K | — | Held |
| Y-Intercept (Hong Kong) Ltd | 34.1K | $311.8K | — | Held |
| FEDERATED HERMES, INC. | 34.1K | $311K | — | Held |
| INTECH INVESTMENT MANAGEMENT LLC | 33.6K | $306.7K | — | Held |
| Calamos Advisors LLC | 32K | $292K | — | Held |
| Gotham Asset Management, LLC | 31.3K | $285.7K | — | Held |
| JANUS HENDERSON GROUP PLC | 29.3K | $268K | — | Held |
| Susquehanna Portfolio Strategies, LLC | 28K | $255.9K | — | Held |
| DIVERSIFIED TRUST CO | 27.8K | $253.8K | — | Held |
| FOX RUN MANAGEMENT, L.L.C. | 25.7K | $234.6K | — | Held |
| NEW YORK STATE COMMON RETIREMENT FUND | 25.5K | $232.8K | — | Held |
| Entropy Technologies, LP | 25.1K | $229.4K | — | Held |
| Quantinno Capital Management LP | 25K | $228.2K | — | Held |
| Police & Firemen's Retirement System of New Jersey | 22.3K | $203.4K | — | Held |
| XTX Topco Ltd | 21.3K | $194.8K | — | Held |
| JONES FINANCIAL COMPANIES LLLP | 20.7K | $189.3K | — | Held |
| State of Wyoming | 20.5K | $187.4K | — | Held |
| LOS ANGELES CAPITAL MANAGEMENT LLC | 11.5K | $178K | — | Held |
| BNP PARIBAS FINANCIAL MARKETS | 19K | $173.5K | — | Held |
| ARIZONA STATE RETIREMENT SYSTEM | 18.5K | $169.3K | — | Held |
| Schonfeld Strategic Advisors LLC | 16.9K | $154.2K | — | Held |
| Tower Research Capital LLC (TRC) | 16.8K | $153K | — | Held |
| Caprock Group, LLC | 16.6K | $151.9K | — | Held |
| Pathstone Holdings, LLC | 16K | $146.1K | — | Held |
| ProShare Advisors LLC | 15.8K | $144.7K | — | Held |
| Walleye Capital LLC | 15.4K | $140.5K | — | Held |
| VOYA INVESTMENT MANAGEMENT LLC | 14.7K | $134K | — | Held |
| KLP KAPITALFORVALTNING AS | 14K | $127.8K | — | Held |
| TWO SIGMA SECURITIES, LLC | 13.8K | $125.9K | — | Held |
| Zurcher Kantonalbank (Zurich Cantonalbank) | 12.7K | $116.1K | — | Held |
| PRELUDE CAPITAL MANAGEMENT, LLC | 11.9K | $108.7K | — | Held |
| RAYMOND JAMES FINANCIAL INC | 11.9K | $108.5K | — | Held |
| SummitTX Capital, L.P. | 11.8K | $107.6K | — | Held |
| Alpine Global Management, LLC | 11.6K | $106.2K | — | Held |
| nVerses Capital, LLC | 11.1K | $101.6K | — | Held |
| GAMMA Investing LLC | 10.9K | $100K | — | Held |
| Pictet Asset Management Holding SA | 10.9K | $99.6K | — | Held |
| Marble Harbor Investment Counsel, LLC | 10.6K | $96.8K | — | Held |
| MILLENNIUM MANAGEMENT LLC | 10.2K | $93.4K | — | Held |
| Olympiad Research LP | 10.1K | $92.3K | — | Held |
| RIK SAYLOR FINANCIAL, INC. | 10K | $91.3K | — | Held |
| OSAIC HOLDINGS, INC. | 7.24K | $66.1K | — | Held |
| Ameritas Investment Partners, Inc. | 6.73K | $61.5K | — | Held |
| AlphaQuest LLC | 6.09K | $55.6K | — | Held |
| Jefferies Financial Group Inc. | 5.64K | $51.5K | — | Held |
| Kemnay Advisory Services Inc. | 4.15K | $37.9K | — | Held |
| MAI Capital Management | 3.27K | $29.9K | — | Held |
| NORTHWESTERN MUTUAL WEALTH MANAGEMENT CO | 3.11K | $28.4K | — | Held |
| Aster Capital Management (DIFC) Ltd | 3K | $27.4K | — | Held |
| Osbon Capital Management LLC | 2.69K | $24.6K | — | Held |
| Quarry LP | 2.63K | $24K | — | Held |
| CITIGROUP INC | 2.35K | $21.4K | — | Held |
| Sterling Capital Management LLC | 1.51K | $13.8K | — | Held |
| Parallel Advisors, LLC | 1.23K | $11.2K | — | Held |
| Rockefeller Capital Management L.P. | 1.21K | $11.1K | — | Held |
| Westfuller Advisors, LLC | 1.2K | $11K | — | Held |
| PNC Financial Services Group, Inc. | 1.18K | $10.7K | — | Held |
| SJS Investment Consulting Inc. | 932 | $8.51K | — | Held |
| NISA INVESTMENT ADVISORS, LLC | 775 | $7.08K | — | Held |
| Optiver Holding B.V. | 740 | $6.76K | — | Held |
| THOMPSON SIEGEL & WALMSLEY LLC | 644.7K | $5.89K | — | Held |
| Essential Partners LLC | 529 | $4.83K | −$4.05K | Cut−$9.87K |
| Manchester Capital Management LLC | 515 | $4.7K | — | Held |
| Rachor Investment Advisory Services, LLC | 513 | $4.68K | — | Held |
| SBI Securities Co., Ltd. | 455 | $4.15K | — | Held |
| Larson Financial Group LLC | 442 | $4.04K | — | Held |
| EverSource Wealth Advisors, LLC | 396 | $3.62K | — | Held |
| HARBOR CAPITAL ADVISORS, INC. | 388.3K | $3.54K | — | Held |
| HRT FINANCIAL LP | 264.3K | $2.41K | — | Held |
| SG Americas Securities, LLC | 76.4K | $2.08K | — | Held |
| Torren Management, LLC | 228 | $2.08K | — | Held |
| Smartleaf Asset Management LLC | 223 | $2.01K | — | Held |
| ACADIAN ASSET MANAGEMENT LLC | 215.8K | $1.97K | — | Held |
| True Wealth Design, LLC | 214 | $1.95K | — | Held |
| US BANCORP \DE\ | 185 | $1.69K | — | Held |
| Versant Capital Management, Inc | 168 | $1.53K | — | Held |
| ROTHSCHILD INVESTMENT LLC | 141 | $1.29K | — | Held |
| Allworth Financial LP | 133 | $1.22K | −$1.02K | Cut−$2.95K |
| Parkside Financial Bank & Trust | 125 | $1.14K | — | Held |
| TEACHERS RETIREMENT SYSTEM OF THE STATE OF KENTUCKY | 119.1K | $1.09K | — | Held |
| Spire Wealth Management | 110 | $1K | — | Held |
| ALGERT GLOBAL LLC | 109.2K | $997 | — | Held |
| BankPlus Trust Department | 100 | $913 | — | Held |
| Bear Mountain Capital, Inc. | 100 | $909 | — | Held |
| GSA CAPITAL PARTNERS LLP | 78.8K | $720 | — | Held |
| Fortitude Family Office, LLC | 55 | $502 | — | Held |
| IFP Advisors, Inc | 47 | $429 | −$61 | Added$295 |
| PRICE T ROWE ASSOCIATES INC /MD/ | 46.5K | $425 | — | Held |
| Sunbelt Securities, Inc. | 40 | $365 | — | Held |
| State of Alaska, Department of Revenue | 38.7K | $353 | — | Held |
| Covestor Ltd | 36.9K | $337 | — | Held |
| Elequin Capital, LP | 20 | $183 | — | Held |
| Farther Finance Advisors, LLC | 20 | $183 | — | Held |
| BNP PARIBAS ASSET MANAGEMENT Holding S.A. | 19.2K | $176 | — | Held |
| Global Retirement Partners, LLC | 19 | $173 | — | Held |
| Keating Financial Advisory Services, Inc. | 11 | $100 | — | Held |
| CWM, LLC | 9.73K | $89 | — | Held |
| CIBC Private Wealth Group LLC | 9 | $82 | — | Held |
| Arax Advisory Partners | 8 | $73 | — | Held |
| AMALGAMATED BANK | 2.72K | $25 | — | Held |