| BML Capital Management, LLC | 5.29M | $6.62M | — | Held |
| ORBIMED ADVISORS LLC | 4.98M | $6.22M | — | Held |
| TANG CAPITAL MANAGEMENT LLC | 4.94M | $6.17M | — | Held |
| ArrowMark Colorado Holdings LLC | 3.09M | $3.86M | — | Held |
| Sio Capital Management, LLC | 2.64M | $3.3M | — | Held |
| Euclidean Capital LLC | 2.63M | $3.28M | — | Held |
| Vanguard | 2.17M | $2.71M | — | Held |
| Stonepine Capital Management, LLC | 1.15M | $1.44M | — | Held |
| BlackRock, Inc. | 1.09M | $1.37M | — | Held |
| TWO SIGMA ADVISERS, LP | 874.1K | $1.09M | — | Held |
| TWO SIGMA INVESTMENTS, LP | 706.1K | $882.6K | — | Held |
| JACOBS LEVY EQUITY MANAGEMENT, INC | 524.4K | $655.4K | — | Held |
| JANE STREET GROUP, LLC | 498.6K | $623.3K | — | Held |
| Aldebaran Capital, LLC | 478K | $597.5K | — | Held |
| MARSHALL WACE, LLP | 465.1K | $581.4K | — | Held |
| GEODE CAPITAL MANAGEMENT, LLC | 446.1K | $557.8K | — | Held |
| BRIDGEWAY CAPITAL MANAGEMENT, LLC | 428.9K | $536.1K | — | Held |
| NORTHERN TRUST CORP | 382.1K | $477.6K | — | Held |
| Walleye Capital LLC | 335.7K | $419.6K | — | Held |
| BANK OF AMERICA CORP /DE/ | 240.1K | $300.2K | −$24.4K | Added$72.7K |
| State Street | 183.2K | $229K | — | Held |
| RBF Capital, LLC | 162.2K | $202.8K | — | Held |
| Assenagon Asset Management S.A. | 140.5K | $175.6K | — | Held |
| EVOKE WEALTH, LLC | 133.1K | $166.4K | — | Held |
| GOLDMAN SACHS GROUP INC | 127K | $158.8K | — | Held |
| Citadel | 118.6K | $148.2K | — | Held |
| XTX Topco Ltd | 104.5K | $130.6K | — | Held |
| ADAR1 Capital Management, LLC | 101K | $126.3K | — | Held |
| MORGAN STANLEY | 98.9K | $123.6K | −$4.86K | Added$78.2K |
| Ikarian Capital, LLC | 89.1K | $111.4K | — | Held |
| SUSQUEHANNA INTERNATIONAL GROUP, LLP | 78.8K | $98.5K | — | Held |
| PNC Financial Services Group, Inc. | 73.2K | $91.5K | — | Held |
| AMERIPRISE FINANCIAL INC | 65.1K | $81.4K | — | Held |
| Seven Fleet Capital Management LP | 63.5K | $79.4K | — | Held |
| Rangeley Capital, LLC | 45.5K | $56.9K | — | Held |
| Velan Capital Investment Management LP | 35K | $43.8K | — | Held |
| HSBC HOLDINGS PLC | 30.9K | $39.4K | — | Held |
| Point72 Asset Management, L.P. | 28.7K | $35.9K | — | Held |
| SIMPLEX TRADING, LLC | 27.9K | $34.8K | — | Held |
| BARCLAYS PLC | 27.5K | $34.4K | −$4.13K | Cut−$4.13K |
| PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO | 27.4K | $34.3K | — | Held |
| BOOTHBAY FUND MANAGEMENT, LLC | 27.1K | $33.9K | — | Held |
| Squarepoint Ops LLC | 26.5K | $33.1K | — | Held |
| TWO SIGMA SECURITIES, LLC | 23.4K | $29.2K | — | Held |
| CITIGROUP INC | 19.3K | $24.1K | — | Held |
| Quadrature Capital Ltd | 18.4K | $23K | — | Held |
| Pathstone Holdings, LLC | 17.5K | $21.9K | — | Held |
| BAILARD, INC. | 17.1K | $21.3K | — | Held |
| NINE MASTS CAPITAL Ltd | 16.6K | $20.8K | — | Held |
| Independent Advisor Alliance | 12.5K | $15.6K | — | Held |
| GTS SECURITIES LLC | 10K | $12.5K | — | Held |
| DEERFIELD MANAGEMENT COMPANY, L.P. | 8.79K | $11K | — | Held |
| Silver Grove Financial Group, Inc. | 5.08K | $6.35K | — | New |
| Torren Management, LLC | 4.72K | $5.89K | — | Held |
| Tower Research Capital LLC (TRC) | 3.9K | $4.88K | — | Held |
| ACADIAN ASSET MANAGEMENT LLC | 2.02M | $2.52K | — | Held |
| UBS Group AG | 1.1K | $1.37K | — | Held |
| JONES FINANCIAL COMPANIES LLLP | 1K | $1.25K | — | Held |
| BNP PARIBAS FINANCIAL MARKETS | 518 | $648 | — | Held |
| JPMORGAN CHASE & CO | 214 | $268 | — | Held |
| ALGERT GLOBAL LLC | 62K | $78 | — | Held |
| Virtu Financial LLC | 12.8K | $16 | — | Held |
| ROYAL BANK OF CANADA | 72 | $0 | — | Held |