| Atlas Venture Life Science Advisors, LLC | 1.14M | $9.11M | — | Held |
| NEA Management Company, LLC | 1.09M | $8.74M | — | Held |
| Lynx1 Capital Management LP | 677.9K | $5.43M | — | Held |
| Nantahala Capital Management, LLC | 600K | $4.81M | — | Held |
| Vanguard | 489.6K | $3.92M | — | Held |
| BlackRock, Inc. | 457.4K | $3.66M | — | Held |
| TCG Crossover Management, LLC | 416.3K | $3.33M | — | Held |
| Siren, L.L.C. | 347.7K | $2.79M | — | Held |
| JANE STREET GROUP, LLC | 288.5K | $2.31M | — | Held |
| 72 Investment Holdings, LLC | 276.8K | $2.22M | — | Held |
| SUSQUEHANNA INTERNATIONAL GROUP, LLP | 196.8K | $1.58M | — | Held |
| GEODE CAPITAL MANAGEMENT, LLC | 162.8K | $1.3M | — | Held |
| MILLENNIUM MANAGEMENT LLC | 161.1K | $1.29M | — | Held |
| State Street | 146.2K | $1.17M | — | Held |
| TWO SIGMA INVESTMENTS, LP | 79K | $632.7K | — | Held |
| Renaissance Technologies | 74.9K | $599.9K | — | Held |
| MORGAN STANLEY | 69.7K | $558.5K | −$2.78M | Cut−$7.46M |
| EMERALD MUTUAL FUND ADVISERS TRUST | 65.5K | $525K | — | Held |
| F/m Investments LLC | 65.5K | $525K | — | Held |
| Citadel | 58.3K | $467.2K | — | Held |
| XTX Topco Ltd | 57.1K | $457.1K | — | Held |
| NORTHERN TRUST CORP | 55.6K | $445.7K | — | Held |
| GOLDMAN SACHS GROUP INC | 52K | $416.1K | — | Held |
| Graham Capital Management, L.P. | 40.8K | $326.5K | — | Held |
| Icahn Capital | 40.6K | $325.6K | — | Held |
| Fidelity | 37.9K | $303.2K | — | Held |
| AIHC Capital Management Ltd | 30.3K | $242.4K | — | Held |
| SIMPLEX TRADING, LLC | 28.2K | $226.3K | — | Held |
| Quantinno Capital Management LP | 27.6K | $221.1K | — | Held |
| Quadrature Capital Ltd | 27.1K | $216.8K | — | Held |
| Engineers Gate Manager LP | 26.9K | $215.3K | — | Held |
| RAYMOND JAMES FINANCIAL INC | 23K | $184.5K | — | Held |
| UBS Group AG | 20K | $160.2K | — | Held |
| MARSHALL WACE, LLP | 19.8K | $158.4K | — | Held |
| Bank of New York Mellon Corp | 18.9K | $151K | — | Held |
| CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 16.3K | $130.8K | — | Held |
| DEUTSCHE BANK AG\ | 12.8K | $102.2K | — | Held |
| TWO SIGMA ADVISERS, LP | 11.2K | $90K | — | Held |
| AMERICAN CENTURY COMPANIES INC | 10.6K | $84.8K | — | Held |
| Jain Global LLC | 10.6K | $84.7K | — | Held |
| GUGGENHEIM CAPITAL LLC | 10.4K | $83.6K | — | Held |
| RHUMBLINE ADVISERS | 10.3K | $82.1K | — | Held |
| AlphaQuest LLC | 7.7K | $61.7K | — | Held |
| WELLS FARGO & COMPANY/MN | 7.52K | $60.3K | — | Held |
| BANK OF AMERICA CORP /DE/ | 5.9K | $47.2K | −$187.6K | Cut−$178K |
| GROUP ONE TRADING LLC | 4K | $32K | — | Held |
| MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd. | 3.15K | $25.2K | — | Held |
| NEW YORK STATE COMMON RETIREMENT FUND | 3K | $24K | — | Held |
| JPMORGAN CHASE & CO | 2.84K | $22.7K | — | Held |
| Russell Investments Group, Ltd. | 1.93K | $15.5K | — | Held |
| North Star Investment Management Corp. | 1.8K | $14.4K | — | Held |
| Steward Partners Investment Advisory, LLC | 1.5K | $12K | — | Held |
| BNP PARIBAS FINANCIAL MARKETS | 1.49K | $11.9K | — | Held |
| OSAIC HOLDINGS, INC. | 1.42K | $11.3K | — | Held |
| Tower Research Capital LLC (TRC) | 1.39K | $11.1K | — | Held |
| Police & Firemen's Retirement System of New Jersey | 1.37K | $11K | — | Held |
| TD Waterhouse Canada Inc. | 800 | $6.4K | — | Held |
| PNC Financial Services Group, Inc. | 740 | $5.93K | — | Held |
| CITIGROUP INC | 607 | $4.86K | — | Held |
| Ameritas Investment Partners, Inc. | 598 | $4.79K | — | Held |
| Legal & General Group Plc | 526 | $4.21K | — | Held |
| LOS ANGELES CAPITAL MANAGEMENT LLC | 110 | $4.01K | — | Held |
| Sterling Capital Management LLC | 458 | $3.67K | — | Held |
| DUNCKER STREETT & CO INC | 437 | $3.5K | — | Held |
| CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM | 408 | $3.27K | — | Held |
| FEDERATED HERMES, INC. | 302 | $2.42K | — | Held |
| ROYAL BANK OF CANADA | 224 | $2K | — | Held |
| Optiver Holding B.V. | 116 | $929 | — | Held |
| GAMMA Investing LLC | 102 | $817 | — | Held |
| UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC | 91 | $729 | — | Held |
| Keating Financial Advisory Services, Inc. | 63 | $505 | — | Held |
| IFP Advisors, Inc | 40 | $320 | −$1.6K | Cut−$1.6K |
| Global Retirement Partners, LLC | 35 | $280 | — | Held |
| ACADIAN ASSET MANAGEMENT LLC | 33.8K | $270 | — | Held |
| ALGERT GLOBAL LLC | 27.1K | $217 | — | Held |
| ROTHSCHILD INVESTMENT LLC | 6 | $48 | — | Held |
| MAI Capital Management | 5 | $41 | — | Held |
| NORTHWESTERN MUTUAL WEALTH MANAGEMENT CO | 2 | $16 | — | Held |
| Allworth Financial LP | 1 | $9 | −$39 | Cut−$4.4K |
| Sunbelt Securities, Inc. | 1 | $8 | — | Held |
| SBI Securities Co., Ltd. | 1 | $8 | — | Held |
| CWM, LLC | 288 | $2 | — | Held |
| AMALGAMATED BANK | 208 | $2 | — | Held |